CD4-binding site alterations in CCR5-using HIV-1 envelopes influencing gp120-CD4 interactions and fusogenicity

CD4-binding site (CD4bs) alterations in gp120 contribute to different pathophysiological phenotypes of CCR5-using (R5) HIV-1 strains, but the potential structural basis is unknown. Here, we characterized functionally diverse R5 envelope (Env) clones (n = 16) to elucidate potential structural alterat...

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Main Authors: Sterjovski, J., Churchill, M., Roche, M., Ellett, A., Farrugia, W., Wesselingh, S., Cunningham, A., Ramsland, Paul, Gorry, P.
Format: Journal Article
Published: 2011
Online Access:http://hdl.handle.net/20.500.11937/6288
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author Sterjovski, J.
Churchill, M.
Roche, M.
Ellett, A.
Farrugia, W.
Wesselingh, S.
Cunningham, A.
Ramsland, Paul
Gorry, P.
author_facet Sterjovski, J.
Churchill, M.
Roche, M.
Ellett, A.
Farrugia, W.
Wesselingh, S.
Cunningham, A.
Ramsland, Paul
Gorry, P.
author_sort Sterjovski, J.
building Curtin Institutional Repository
collection Online Access
description CD4-binding site (CD4bs) alterations in gp120 contribute to different pathophysiological phenotypes of CCR5-using (R5) HIV-1 strains, but the potential structural basis is unknown. Here, we characterized functionally diverse R5 envelope (Env) clones (n = 16) to elucidate potential structural alterations within the gp120 CD4bs that influence Env function. Initially, we showed that the magnitude of gp120-CD4-binding correlates with increased fusogenicity and reduced CD4 dependence. Analysis of three-dimensional gp120 structural models revealed two CD4bs variants, D279 and N362, that were associated with reduced CD4 dependence. Further structural analysis showed that a wider aperture of the predicted CD4bs cavity, as constrained by the inner-most atoms at the gp120 V1V2 stem and the V5 loop, was associated with amino acid alterations within V5 and correlated with increased gp120-CD4 binding and increased fusogenicity. Our results provide evidence that the gp120 V5 loop may alter CD4bs conformation and contribute to increased gp120-CD4 interactions and Env fusogenicity.
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spelling curtin-20.500.11937-62882017-09-13T14:41:37Z CD4-binding site alterations in CCR5-using HIV-1 envelopes influencing gp120-CD4 interactions and fusogenicity Sterjovski, J. Churchill, M. Roche, M. Ellett, A. Farrugia, W. Wesselingh, S. Cunningham, A. Ramsland, Paul Gorry, P. CD4-binding site (CD4bs) alterations in gp120 contribute to different pathophysiological phenotypes of CCR5-using (R5) HIV-1 strains, but the potential structural basis is unknown. Here, we characterized functionally diverse R5 envelope (Env) clones (n = 16) to elucidate potential structural alterations within the gp120 CD4bs that influence Env function. Initially, we showed that the magnitude of gp120-CD4-binding correlates with increased fusogenicity and reduced CD4 dependence. Analysis of three-dimensional gp120 structural models revealed two CD4bs variants, D279 and N362, that were associated with reduced CD4 dependence. Further structural analysis showed that a wider aperture of the predicted CD4bs cavity, as constrained by the inner-most atoms at the gp120 V1V2 stem and the V5 loop, was associated with amino acid alterations within V5 and correlated with increased gp120-CD4 binding and increased fusogenicity. Our results provide evidence that the gp120 V5 loop may alter CD4bs conformation and contribute to increased gp120-CD4 interactions and Env fusogenicity. 2011 Journal Article http://hdl.handle.net/20.500.11937/6288 10.1016/j.virol.2010.12.010 unknown
spellingShingle Sterjovski, J.
Churchill, M.
Roche, M.
Ellett, A.
Farrugia, W.
Wesselingh, S.
Cunningham, A.
Ramsland, Paul
Gorry, P.
CD4-binding site alterations in CCR5-using HIV-1 envelopes influencing gp120-CD4 interactions and fusogenicity
title CD4-binding site alterations in CCR5-using HIV-1 envelopes influencing gp120-CD4 interactions and fusogenicity
title_full CD4-binding site alterations in CCR5-using HIV-1 envelopes influencing gp120-CD4 interactions and fusogenicity
title_fullStr CD4-binding site alterations in CCR5-using HIV-1 envelopes influencing gp120-CD4 interactions and fusogenicity
title_full_unstemmed CD4-binding site alterations in CCR5-using HIV-1 envelopes influencing gp120-CD4 interactions and fusogenicity
title_short CD4-binding site alterations in CCR5-using HIV-1 envelopes influencing gp120-CD4 interactions and fusogenicity
title_sort cd4-binding site alterations in ccr5-using hiv-1 envelopes influencing gp120-cd4 interactions and fusogenicity
url http://hdl.handle.net/20.500.11937/6288