mTORC1 activity repression by late endosomal phosphatidylinositol 3,4-bisphosphate

Nutrient sensing by mechanistic target of rapamycin complex 1 (mTORC1) on lysosomes and late endosomes (LyLEs) regulates cell growth. Many factors stimulate mTORC1 activity, including the production of phosphatidylinositol 3,4,5-trisphosphate [PI(3,4,5)P3] by class I phosphatidylinositol 3-kinases (...

Full description

Bibliographic Details
Main Authors: Marat, A., Wallroth, A., Lo, W., Müller, R., Norata, G., Falasca, Marco, Schultz, C., Haucke, V.
Format: Journal Article
Published: The American Association for the Advancement of Science 2017
Online Access:http://hdl.handle.net/20.500.11937/54003
_version_ 1848759280529833984
author Marat, A.
Wallroth, A.
Lo, W.
Müller, R.
Norata, G.
Falasca, Marco
Schultz, C.
Haucke, V.
author_facet Marat, A.
Wallroth, A.
Lo, W.
Müller, R.
Norata, G.
Falasca, Marco
Schultz, C.
Haucke, V.
author_sort Marat, A.
building Curtin Institutional Repository
collection Online Access
description Nutrient sensing by mechanistic target of rapamycin complex 1 (mTORC1) on lysosomes and late endosomes (LyLEs) regulates cell growth. Many factors stimulate mTORC1 activity, including the production of phosphatidylinositol 3,4,5-trisphosphate [PI(3,4,5)P3] by class I phosphatidylinositol 3-kinases (PI3Ks) at the plasma membrane. We investigated mechanisms that repress mTORC1 under conditions of growth factor deprivation. We identified phosphatidylinositol 3,4-bisphosphate [PI(3,4)P2], synthesized by class II PI3K b (PI3KC2b) at LyLEs, as a negative regulator of mTORC1, whereas loss of PI3KC2b hyperactivated mTORC1. Growth factor deprivation induced the association of PI3KC2b with the Raptor subunit of mTORC1. Local PI(3,4)P2 synthesis triggered repression of mTORC1 activity through association of Raptor with inhibitory 14-3-3 proteins. These results unravel an unexpected function for local PI(3,4)P2 production in shutting off mTORC1.
first_indexed 2025-11-14T09:57:22Z
format Journal Article
id curtin-20.500.11937-54003
institution Curtin University Malaysia
institution_category Local University
last_indexed 2025-11-14T09:57:22Z
publishDate 2017
publisher The American Association for the Advancement of Science
recordtype eprints
repository_type Digital Repository
spelling curtin-20.500.11937-540032017-09-13T15:46:23Z mTORC1 activity repression by late endosomal phosphatidylinositol 3,4-bisphosphate Marat, A. Wallroth, A. Lo, W. Müller, R. Norata, G. Falasca, Marco Schultz, C. Haucke, V. Nutrient sensing by mechanistic target of rapamycin complex 1 (mTORC1) on lysosomes and late endosomes (LyLEs) regulates cell growth. Many factors stimulate mTORC1 activity, including the production of phosphatidylinositol 3,4,5-trisphosphate [PI(3,4,5)P3] by class I phosphatidylinositol 3-kinases (PI3Ks) at the plasma membrane. We investigated mechanisms that repress mTORC1 under conditions of growth factor deprivation. We identified phosphatidylinositol 3,4-bisphosphate [PI(3,4)P2], synthesized by class II PI3K b (PI3KC2b) at LyLEs, as a negative regulator of mTORC1, whereas loss of PI3KC2b hyperactivated mTORC1. Growth factor deprivation induced the association of PI3KC2b with the Raptor subunit of mTORC1. Local PI(3,4)P2 synthesis triggered repression of mTORC1 activity through association of Raptor with inhibitory 14-3-3 proteins. These results unravel an unexpected function for local PI(3,4)P2 production in shutting off mTORC1. 2017 Journal Article http://hdl.handle.net/20.500.11937/54003 10.1126/science.aaf8310 The American Association for the Advancement of Science restricted
spellingShingle Marat, A.
Wallroth, A.
Lo, W.
Müller, R.
Norata, G.
Falasca, Marco
Schultz, C.
Haucke, V.
mTORC1 activity repression by late endosomal phosphatidylinositol 3,4-bisphosphate
title mTORC1 activity repression by late endosomal phosphatidylinositol 3,4-bisphosphate
title_full mTORC1 activity repression by late endosomal phosphatidylinositol 3,4-bisphosphate
title_fullStr mTORC1 activity repression by late endosomal phosphatidylinositol 3,4-bisphosphate
title_full_unstemmed mTORC1 activity repression by late endosomal phosphatidylinositol 3,4-bisphosphate
title_short mTORC1 activity repression by late endosomal phosphatidylinositol 3,4-bisphosphate
title_sort mtorc1 activity repression by late endosomal phosphatidylinositol 3,4-bisphosphate
url http://hdl.handle.net/20.500.11937/54003