AUX1-mediated root hair auxin influx governs SCFTIR1/AFB-type Ca2+ signaling

Auxin is a key regulator of plant growth and development, but the causal relationship between hormone transport and root responses remains unresolved. Here we describe auxin uptake, together with early steps in signaling, in Arabidopsis root hairs. Using intracellular microelectrodes we show membran...

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Main Authors: Dindas, Julian, Scherzer, Sönke, Roelfsema, M. Rob G., von Meyer, Katharina, Müller, Heike M., Al-Rasheid, K.A.S., Palme, Klaus, Dietrich, Petra, Becker, Dirk, Bennett, Malcolm J., Hedrich, Rainer
Format: Article
Language:English
Published: Nature Publishing Group 2018
Online Access:https://eprints.nottingham.ac.uk/52988/
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author Dindas, Julian
Scherzer, Sönke
Roelfsema, M. Rob G.
von Meyer, Katharina
Müller, Heike M.
Al-Rasheid, K.A.S.
Palme, Klaus
Dietrich, Petra
Becker, Dirk
Bennett, Malcolm J.
Hedrich, Rainer
author_facet Dindas, Julian
Scherzer, Sönke
Roelfsema, M. Rob G.
von Meyer, Katharina
Müller, Heike M.
Al-Rasheid, K.A.S.
Palme, Klaus
Dietrich, Petra
Becker, Dirk
Bennett, Malcolm J.
Hedrich, Rainer
author_sort Dindas, Julian
building Nottingham Research Data Repository
collection Online Access
description Auxin is a key regulator of plant growth and development, but the causal relationship between hormone transport and root responses remains unresolved. Here we describe auxin uptake, together with early steps in signaling, in Arabidopsis root hairs. Using intracellular microelectrodes we show membrane depolarization, in response to IAA in a concentration- and pH-dependent manner. This depolarization is strongly impaired in aux1 mutants, indicating that AUX1 is the major transporter for auxin uptake in root hairs. Local intracellular auxin application triggers Ca2+ signals that propagate as long-distance waves between root cells and modulate their auxin responses. AUX1-mediated IAA transport, as well as IAA- triggered calcium signals, are blocked by treatment with the SCFTIR1/AFB - inhibitor auxinole. Further, they are strongly reduced in the tir1afb2afb3 and the cngc14 mutant. Our study reveals that the AUX1 transporter, the SCFTIR1/AFB receptor and the CNGC14 Ca2+ channel, mediate fast auxin signaling in roots.
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spelling nottingham-529882020-05-08T08:02:33Z https://eprints.nottingham.ac.uk/52988/ AUX1-mediated root hair auxin influx governs SCFTIR1/AFB-type Ca2+ signaling Dindas, Julian Scherzer, Sönke Roelfsema, M. Rob G. von Meyer, Katharina Müller, Heike M. Al-Rasheid, K.A.S. Palme, Klaus Dietrich, Petra Becker, Dirk Bennett, Malcolm J. Hedrich, Rainer Auxin is a key regulator of plant growth and development, but the causal relationship between hormone transport and root responses remains unresolved. Here we describe auxin uptake, together with early steps in signaling, in Arabidopsis root hairs. Using intracellular microelectrodes we show membrane depolarization, in response to IAA in a concentration- and pH-dependent manner. This depolarization is strongly impaired in aux1 mutants, indicating that AUX1 is the major transporter for auxin uptake in root hairs. Local intracellular auxin application triggers Ca2+ signals that propagate as long-distance waves between root cells and modulate their auxin responses. AUX1-mediated IAA transport, as well as IAA- triggered calcium signals, are blocked by treatment with the SCFTIR1/AFB - inhibitor auxinole. Further, they are strongly reduced in the tir1afb2afb3 and the cngc14 mutant. Our study reveals that the AUX1 transporter, the SCFTIR1/AFB receptor and the CNGC14 Ca2+ channel, mediate fast auxin signaling in roots. Nature Publishing Group 2018-03-21 Article PeerReviewed application/pdf en cc_by https://eprints.nottingham.ac.uk/52988/1/s41467-018-03582-5.pdf Dindas, Julian, Scherzer, Sönke, Roelfsema, M. Rob G., von Meyer, Katharina, Müller, Heike M., Al-Rasheid, K.A.S., Palme, Klaus, Dietrich, Petra, Becker, Dirk, Bennett, Malcolm J. and Hedrich, Rainer (2018) AUX1-mediated root hair auxin influx governs SCFTIR1/AFB-type Ca2+ signaling. Nature Communications, 9 (1). 1174/1-1174/10. ISSN 2041-1723 https://www.nature.com/articles/s41467-018-03582-5 doi:10.1038/s41467-018-03582-5 doi:10.1038/s41467-018-03582-5
spellingShingle Dindas, Julian
Scherzer, Sönke
Roelfsema, M. Rob G.
von Meyer, Katharina
Müller, Heike M.
Al-Rasheid, K.A.S.
Palme, Klaus
Dietrich, Petra
Becker, Dirk
Bennett, Malcolm J.
Hedrich, Rainer
AUX1-mediated root hair auxin influx governs SCFTIR1/AFB-type Ca2+ signaling
title AUX1-mediated root hair auxin influx governs SCFTIR1/AFB-type Ca2+ signaling
title_full AUX1-mediated root hair auxin influx governs SCFTIR1/AFB-type Ca2+ signaling
title_fullStr AUX1-mediated root hair auxin influx governs SCFTIR1/AFB-type Ca2+ signaling
title_full_unstemmed AUX1-mediated root hair auxin influx governs SCFTIR1/AFB-type Ca2+ signaling
title_short AUX1-mediated root hair auxin influx governs SCFTIR1/AFB-type Ca2+ signaling
title_sort aux1-mediated root hair auxin influx governs scftir1/afb-type ca2+ signaling
url https://eprints.nottingham.ac.uk/52988/
https://eprints.nottingham.ac.uk/52988/
https://eprints.nottingham.ac.uk/52988/