Flexible theta sequence compression mediated via phase precessing interneurons

Encoding of behavioral episodes as spike sequences during hippocampal theta oscillations provides a neural substrate for computations on events extended across time and space. However, the mechanisms underlying the numerous and diverse experimentally observed properties of theta sequences remain poo...

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Main Authors: Chadwick, Angus, van Rossum, Mark C.W., Nolan, Matthew F.
Format: Article
Published: eLife Sciences Publications 2016
Online Access:https://eprints.nottingham.ac.uk/49616/
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author Chadwick, Angus
van Rossum, Mark C.W.
Nolan, Matthew F.
author_facet Chadwick, Angus
van Rossum, Mark C.W.
Nolan, Matthew F.
author_sort Chadwick, Angus
building Nottingham Research Data Repository
collection Online Access
description Encoding of behavioral episodes as spike sequences during hippocampal theta oscillations provides a neural substrate for computations on events extended across time and space. However, the mechanisms underlying the numerous and diverse experimentally observed properties of theta sequences remain poorly understood. Here we account for theta sequences using a novel model constrained by the septo-hippocampal circuitry. We show that when spontaneously active interneurons integrate spatial signals and theta frequency pacemaker inputs, they generate phase precessing action potentials that can coordinate theta sequences in place cell populations. We reveal novel constraints on sequence generation, predict cellular properties and neural dynamics that characterize sequence compression, identify circuit organization principles for high capacity sequential representation, and show that theta sequences can be used as substrates for association of conditioned stimuli with recent and upcoming events. Our results suggest mechanisms for flexible sequence compression that are suited to associative learning across an animal's lifespan.
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spelling nottingham-496162020-05-04T18:26:53Z https://eprints.nottingham.ac.uk/49616/ Flexible theta sequence compression mediated via phase precessing interneurons Chadwick, Angus van Rossum, Mark C.W. Nolan, Matthew F. Encoding of behavioral episodes as spike sequences during hippocampal theta oscillations provides a neural substrate for computations on events extended across time and space. However, the mechanisms underlying the numerous and diverse experimentally observed properties of theta sequences remain poorly understood. Here we account for theta sequences using a novel model constrained by the septo-hippocampal circuitry. We show that when spontaneously active interneurons integrate spatial signals and theta frequency pacemaker inputs, they generate phase precessing action potentials that can coordinate theta sequences in place cell populations. We reveal novel constraints on sequence generation, predict cellular properties and neural dynamics that characterize sequence compression, identify circuit organization principles for high capacity sequential representation, and show that theta sequences can be used as substrates for association of conditioned stimuli with recent and upcoming events. Our results suggest mechanisms for flexible sequence compression that are suited to associative learning across an animal's lifespan. eLife Sciences Publications 2016-12-08 Article PeerReviewed Chadwick, Angus, van Rossum, Mark C.W. and Nolan, Matthew F. (2016) Flexible theta sequence compression mediated via phase precessing interneurons. eLife, 5 . e20349/1-e20349/30. ISSN 2050-084X https://elifesciences.org/articles/20349 doi:10.7554/eLife.20349 doi:10.7554/eLife.20349
spellingShingle Chadwick, Angus
van Rossum, Mark C.W.
Nolan, Matthew F.
Flexible theta sequence compression mediated via phase precessing interneurons
title Flexible theta sequence compression mediated via phase precessing interneurons
title_full Flexible theta sequence compression mediated via phase precessing interneurons
title_fullStr Flexible theta sequence compression mediated via phase precessing interneurons
title_full_unstemmed Flexible theta sequence compression mediated via phase precessing interneurons
title_short Flexible theta sequence compression mediated via phase precessing interneurons
title_sort flexible theta sequence compression mediated via phase precessing interneurons
url https://eprints.nottingham.ac.uk/49616/
https://eprints.nottingham.ac.uk/49616/
https://eprints.nottingham.ac.uk/49616/