Applying asymptotic methods to synthetic biology: modelling the reaction kinetics of the mevalonate pathway

The mevalonate pathway is normally found in eukaryotes, and allows for the production of isoprenoids, a useful class of organic compounds. This pathway has been successfully introduced to Escherichia coli, enabling a biosynthetic production route for many isoprenoids. In this paper, we develop and s...

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Main Authors: Dalwadi, Mohit P., Garavaglia, Marco, Webb, Joseph P., King, John R., Minton, Nigel P.
Format: Article
Published: Elsevier 2018
Subjects:
Online Access:https://eprints.nottingham.ac.uk/48475/
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author Dalwadi, Mohit P.
Garavaglia, Marco
Webb, Joseph P.
King, John R.
Minton, Nigel P.
author_facet Dalwadi, Mohit P.
Garavaglia, Marco
Webb, Joseph P.
King, John R.
Minton, Nigel P.
author_sort Dalwadi, Mohit P.
building Nottingham Research Data Repository
collection Online Access
description The mevalonate pathway is normally found in eukaryotes, and allows for the production of isoprenoids, a useful class of organic compounds. This pathway has been successfully introduced to Escherichia coli, enabling a biosynthetic production route for many isoprenoids. In this paper, we develop and solve a mathematical model for the concentration of metabolites in the mevalonate pathway over time, accounting for the loss of acetyl-CoA to other metabolic pathways. Additionally, we successfully test our theoretical predictions experimentally by introducing part of the pathway into Cupriavidus necator. In our model, we exploit the natural separation of time scales as well as of metabolite concentrations to make significant asymptotic progress in understanding the system. We confirm that our asymptotic results agree well with numerical simulations, the former enabling us to predict the most important reactions to increase isopentenyl diphosphate production whilst minimizing the levels of HMG-CoA, which inhibits cell growth. Thus, our mathematical model allows us to recommend the upregulation of certain combinations of enzymes to improve production through the mevalonate pathway.
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spelling nottingham-484752020-05-04T19:32:01Z https://eprints.nottingham.ac.uk/48475/ Applying asymptotic methods to synthetic biology: modelling the reaction kinetics of the mevalonate pathway Dalwadi, Mohit P. Garavaglia, Marco Webb, Joseph P. King, John R. Minton, Nigel P. The mevalonate pathway is normally found in eukaryotes, and allows for the production of isoprenoids, a useful class of organic compounds. This pathway has been successfully introduced to Escherichia coli, enabling a biosynthetic production route for many isoprenoids. In this paper, we develop and solve a mathematical model for the concentration of metabolites in the mevalonate pathway over time, accounting for the loss of acetyl-CoA to other metabolic pathways. Additionally, we successfully test our theoretical predictions experimentally by introducing part of the pathway into Cupriavidus necator. In our model, we exploit the natural separation of time scales as well as of metabolite concentrations to make significant asymptotic progress in understanding the system. We confirm that our asymptotic results agree well with numerical simulations, the former enabling us to predict the most important reactions to increase isopentenyl diphosphate production whilst minimizing the levels of HMG-CoA, which inhibits cell growth. Thus, our mathematical model allows us to recommend the upregulation of certain combinations of enzymes to improve production through the mevalonate pathway. Elsevier 2018-02-14 Article PeerReviewed Dalwadi, Mohit P., Garavaglia, Marco, Webb, Joseph P., King, John R. and Minton, Nigel P. (2018) Applying asymptotic methods to synthetic biology: modelling the reaction kinetics of the mevalonate pathway. Journal of Theoretical Biology, 439 . pp. 39-49. ISSN 1095-8541 asymptotic analysis; metabolic pathways; isoprenoid production https://www.sciencedirect.com/science/article/pii/S0022519317305295 doi:10.1016/j.jtbi.2017.11.022 doi:10.1016/j.jtbi.2017.11.022
spellingShingle asymptotic analysis; metabolic pathways; isoprenoid production
Dalwadi, Mohit P.
Garavaglia, Marco
Webb, Joseph P.
King, John R.
Minton, Nigel P.
Applying asymptotic methods to synthetic biology: modelling the reaction kinetics of the mevalonate pathway
title Applying asymptotic methods to synthetic biology: modelling the reaction kinetics of the mevalonate pathway
title_full Applying asymptotic methods to synthetic biology: modelling the reaction kinetics of the mevalonate pathway
title_fullStr Applying asymptotic methods to synthetic biology: modelling the reaction kinetics of the mevalonate pathway
title_full_unstemmed Applying asymptotic methods to synthetic biology: modelling the reaction kinetics of the mevalonate pathway
title_short Applying asymptotic methods to synthetic biology: modelling the reaction kinetics of the mevalonate pathway
title_sort applying asymptotic methods to synthetic biology: modelling the reaction kinetics of the mevalonate pathway
topic asymptotic analysis; metabolic pathways; isoprenoid production
url https://eprints.nottingham.ac.uk/48475/
https://eprints.nottingham.ac.uk/48475/
https://eprints.nottingham.ac.uk/48475/