Dynamic regulation of auxin response during rice development revealed by newly established hormone biosensor markers

The hormone auxin is critical for many plant developmental processes. Unlike the model eudicot plant Arabidopsis (Arabidopsis thaliana), auxin distribution and signaling in rice tissues has not been systematically investigated due to the absence of suitable auxin response reporters. In this study we...

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Main Authors: Yang, Jing, Yuan, Zheng, Meng, Qingcai, Huang, Guoqiang, Périn, Christophe, Bureau, Charlotte, Meunier, Anne-Cécile, Ingouff, Mathieu, Bennett, Malcolm J., Liang, Wanqi, Zhang, Dabing
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Published: Frontiers Media 2017
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Online Access:https://eprints.nottingham.ac.uk/44429/
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author Yang, Jing
Yuan, Zheng
Meng, Qingcai
Huang, Guoqiang
Périn, Christophe
Bureau, Charlotte
Meunier, Anne-Cécile
Ingouff, Mathieu
Bennett, Malcolm J.
Liang, Wanqi
Zhang, Dabing
author_facet Yang, Jing
Yuan, Zheng
Meng, Qingcai
Huang, Guoqiang
Périn, Christophe
Bureau, Charlotte
Meunier, Anne-Cécile
Ingouff, Mathieu
Bennett, Malcolm J.
Liang, Wanqi
Zhang, Dabing
author_sort Yang, Jing
building Nottingham Research Data Repository
collection Online Access
description The hormone auxin is critical for many plant developmental processes. Unlike the model eudicot plant Arabidopsis (Arabidopsis thaliana), auxin distribution and signaling in rice tissues has not been systematically investigated due to the absence of suitable auxin response reporters. In this study we observed the conservation of auxin signalling components between Arabidopsis and model monocot crop rice (Oryza sativa), and generated complementary types of auxin biosensor constructs, one derived from the Aux/IAA-based biosensor DII-VENUS but constitutively driven by maize ubiquitin-1 promoter, and the other termed DR5-VENUS in which a synthetic auxin-responsive promoter (DR5rev) was used to drive expression of the yellow fluorescent protein (YFP). Using the obtained transgenic lines, we observed that during the vegetative development, accumulation of DR5-VENUS signal was at young and mature leaves, tiller buds and stem base. Notably, abundant DR5-VENUS signals were observed in the cytoplasm of cortex cells surrounding lateral root primordia (LRP) in rice. In addition, auxinmaxima and dynamic re-localization were seen at the initiation sites of inflorescence and spikelet primordia including branch meristems (BMs), female and male organs. The comparison of these observations among Arabidopsis, rice and maize suggests the unique role of auxin in regulating rice lateral root emergence and reproduction. Moreover, protein localization of auxin transporters PIN1 homologs and GFP tagged OsAUX1 overlapped with DR5-VENUS during spikelet development, helping validate these auxin response reporters are reliable markers in rice. This work firstly reveals the direct correspondence between auxin distribution and rice reproductive and root development at tissue and cellular level, and provides high-resolution auxin tools to probe fundamental developmental processes in rice and to establish links between auxin, development and agronomical traits like yield or root architecture.
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spelling nottingham-444292020-05-04T18:36:46Z https://eprints.nottingham.ac.uk/44429/ Dynamic regulation of auxin response during rice development revealed by newly established hormone biosensor markers Yang, Jing Yuan, Zheng Meng, Qingcai Huang, Guoqiang Périn, Christophe Bureau, Charlotte Meunier, Anne-Cécile Ingouff, Mathieu Bennett, Malcolm J. Liang, Wanqi Zhang, Dabing The hormone auxin is critical for many plant developmental processes. Unlike the model eudicot plant Arabidopsis (Arabidopsis thaliana), auxin distribution and signaling in rice tissues has not been systematically investigated due to the absence of suitable auxin response reporters. In this study we observed the conservation of auxin signalling components between Arabidopsis and model monocot crop rice (Oryza sativa), and generated complementary types of auxin biosensor constructs, one derived from the Aux/IAA-based biosensor DII-VENUS but constitutively driven by maize ubiquitin-1 promoter, and the other termed DR5-VENUS in which a synthetic auxin-responsive promoter (DR5rev) was used to drive expression of the yellow fluorescent protein (YFP). Using the obtained transgenic lines, we observed that during the vegetative development, accumulation of DR5-VENUS signal was at young and mature leaves, tiller buds and stem base. Notably, abundant DR5-VENUS signals were observed in the cytoplasm of cortex cells surrounding lateral root primordia (LRP) in rice. In addition, auxinmaxima and dynamic re-localization were seen at the initiation sites of inflorescence and spikelet primordia including branch meristems (BMs), female and male organs. The comparison of these observations among Arabidopsis, rice and maize suggests the unique role of auxin in regulating rice lateral root emergence and reproduction. Moreover, protein localization of auxin transporters PIN1 homologs and GFP tagged OsAUX1 overlapped with DR5-VENUS during spikelet development, helping validate these auxin response reporters are reliable markers in rice. This work firstly reveals the direct correspondence between auxin distribution and rice reproductive and root development at tissue and cellular level, and provides high-resolution auxin tools to probe fundamental developmental processes in rice and to establish links between auxin, development and agronomical traits like yield or root architecture. Frontiers Media 2017-03-07 Article PeerReviewed Yang, Jing, Yuan, Zheng, Meng, Qingcai, Huang, Guoqiang, Périn, Christophe, Bureau, Charlotte, Meunier, Anne-Cécile, Ingouff, Mathieu, Bennett, Malcolm J., Liang, Wanqi and Zhang, Dabing (2017) Dynamic regulation of auxin response during rice development revealed by newly established hormone biosensor markers. Frontiers in Plant Science, 8 . p. 256. ISSN 1664-462X rice auxin reporter lateral root formation inflorescence spikelet meristem http://journal.frontiersin.org/article/10.3389/fpls.2017.00256/full doi:10.3389/fpls.2017.00256 doi:10.3389/fpls.2017.00256
spellingShingle rice
auxin
reporter
lateral root formation
inflorescence
spikelet
meristem
Yang, Jing
Yuan, Zheng
Meng, Qingcai
Huang, Guoqiang
Périn, Christophe
Bureau, Charlotte
Meunier, Anne-Cécile
Ingouff, Mathieu
Bennett, Malcolm J.
Liang, Wanqi
Zhang, Dabing
Dynamic regulation of auxin response during rice development revealed by newly established hormone biosensor markers
title Dynamic regulation of auxin response during rice development revealed by newly established hormone biosensor markers
title_full Dynamic regulation of auxin response during rice development revealed by newly established hormone biosensor markers
title_fullStr Dynamic regulation of auxin response during rice development revealed by newly established hormone biosensor markers
title_full_unstemmed Dynamic regulation of auxin response during rice development revealed by newly established hormone biosensor markers
title_short Dynamic regulation of auxin response during rice development revealed by newly established hormone biosensor markers
title_sort dynamic regulation of auxin response during rice development revealed by newly established hormone biosensor markers
topic rice
auxin
reporter
lateral root formation
inflorescence
spikelet
meristem
url https://eprints.nottingham.ac.uk/44429/
https://eprints.nottingham.ac.uk/44429/
https://eprints.nottingham.ac.uk/44429/