Hypothalamic over-expression of VGF in the Siberian hamster increases energy expenditure and reduces body weight gain

VGF (non-acronymic) was first highlighted to have a role in energy homeostasis through experiments involving dietary manipulation in mice. Fasting increased VGF mRNA in the Arc and levels were subsequently reduced upon refeeding. This anabolic role for VGF was supported by observations in a VGF null...

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Main Authors: Lewis, Jo E., Brameld, John M., Hill, Phil, Cocco, Cristina, Noli, Barbara, Ferri, Gian-Luca, Barrett, Perry, Ebling, Francis J.P., Jethwa, P.H.
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Published: Public Library of Science 2017
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Online Access:https://eprints.nottingham.ac.uk/40849/
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author Lewis, Jo E.
Brameld, John M.
Hill, Phil
Cocco, Cristina
Noli, Barbara
Ferri, Gian-Luca
Barrett, Perry
Ebling, Francis J.P.
Jethwa, P.H.
author_facet Lewis, Jo E.
Brameld, John M.
Hill, Phil
Cocco, Cristina
Noli, Barbara
Ferri, Gian-Luca
Barrett, Perry
Ebling, Francis J.P.
Jethwa, P.H.
author_sort Lewis, Jo E.
building Nottingham Research Data Repository
collection Online Access
description VGF (non-acronymic) was first highlighted to have a role in energy homeostasis through experiments involving dietary manipulation in mice. Fasting increased VGF mRNA in the Arc and levels were subsequently reduced upon refeeding. This anabolic role for VGF was supported by observations in a VGF null (VGF-/-) mouse and in the diet-induced and gold-thioglucose obese mice. However, this anabolic role for VGF has not been supported by a number of subsequent studies investigating the physiological effects of VGF-derived peptides. Intracerebroventricular (ICV) infusion of TLQP-21 increased resting energy expenditure and rectal temperature in mice and protected against diet-induced obesity. Similarly, ICV infusion of TLQP-21 into Siberian hamsters significantly reduced body weight, but this was due to a decrease in food intake, with no effect on energy expenditure. Subsequently NERP-2 was shown to increase food intake in rats via the orexin system, suggesting opposing roles for these VGF-derived peptides. Thus to further elucidate the role of hypothalamic VGF in the regulation of energy homeostasis we utilised a recombinant adeno-associated viral vector to over-express VGF in adult male Siberian hamsters, thus avoiding any developmental effects or associated functional compensation. Initially, hypothalamic over-expression of VGF in adult Siberian hamsters produced no effect on metabolic parameters, but by 12 weeks post-infusion hamsters had increased oxygen consumption and a tendency to increased carbon dioxide production; this attenuated body weight gain, reduced interscapular white adipose tissue and resulted in a compensatory increase in food intake. These observed changes in energy expenditure and food intake were associated with an increase in the hypothalamic contents of the VGF-derived peptides AQEE, TLQP and NERP-2. The complex phenotype of the VGF-/- mice is a likely consequence of global ablation of the gene and its derived peptides during development, as well as in the adult.
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spelling nottingham-408492020-05-04T18:33:38Z https://eprints.nottingham.ac.uk/40849/ Hypothalamic over-expression of VGF in the Siberian hamster increases energy expenditure and reduces body weight gain Lewis, Jo E. Brameld, John M. Hill, Phil Cocco, Cristina Noli, Barbara Ferri, Gian-Luca Barrett, Perry Ebling, Francis J.P. Jethwa, P.H. VGF (non-acronymic) was first highlighted to have a role in energy homeostasis through experiments involving dietary manipulation in mice. Fasting increased VGF mRNA in the Arc and levels were subsequently reduced upon refeeding. This anabolic role for VGF was supported by observations in a VGF null (VGF-/-) mouse and in the diet-induced and gold-thioglucose obese mice. However, this anabolic role for VGF has not been supported by a number of subsequent studies investigating the physiological effects of VGF-derived peptides. Intracerebroventricular (ICV) infusion of TLQP-21 increased resting energy expenditure and rectal temperature in mice and protected against diet-induced obesity. Similarly, ICV infusion of TLQP-21 into Siberian hamsters significantly reduced body weight, but this was due to a decrease in food intake, with no effect on energy expenditure. Subsequently NERP-2 was shown to increase food intake in rats via the orexin system, suggesting opposing roles for these VGF-derived peptides. Thus to further elucidate the role of hypothalamic VGF in the regulation of energy homeostasis we utilised a recombinant adeno-associated viral vector to over-express VGF in adult male Siberian hamsters, thus avoiding any developmental effects or associated functional compensation. Initially, hypothalamic over-expression of VGF in adult Siberian hamsters produced no effect on metabolic parameters, but by 12 weeks post-infusion hamsters had increased oxygen consumption and a tendency to increased carbon dioxide production; this attenuated body weight gain, reduced interscapular white adipose tissue and resulted in a compensatory increase in food intake. These observed changes in energy expenditure and food intake were associated with an increase in the hypothalamic contents of the VGF-derived peptides AQEE, TLQP and NERP-2. The complex phenotype of the VGF-/- mice is a likely consequence of global ablation of the gene and its derived peptides during development, as well as in the adult. Public Library of Science 2017-02-24 Article PeerReviewed Lewis, Jo E., Brameld, John M., Hill, Phil, Cocco, Cristina, Noli, Barbara, Ferri, Gian-Luca, Barrett, Perry, Ebling, Francis J.P. and Jethwa, P.H. (2017) Hypothalamic over-expression of VGF in the Siberian hamster increases energy expenditure and reduces body weight gain. PLoS ONE, 12 . e01727/1-e01727/14. ISSN 1932-6203 Siberian hamsters VGF hypothalamus adeno-associated viral virus http://journals.plos.org/plosone/article?id=10.1371/journal.pone.0172724 doi:10.1371/journal.pone.0172724 doi:10.1371/journal.pone.0172724
spellingShingle Siberian hamsters
VGF
hypothalamus
adeno-associated viral virus
Lewis, Jo E.
Brameld, John M.
Hill, Phil
Cocco, Cristina
Noli, Barbara
Ferri, Gian-Luca
Barrett, Perry
Ebling, Francis J.P.
Jethwa, P.H.
Hypothalamic over-expression of VGF in the Siberian hamster increases energy expenditure and reduces body weight gain
title Hypothalamic over-expression of VGF in the Siberian hamster increases energy expenditure and reduces body weight gain
title_full Hypothalamic over-expression of VGF in the Siberian hamster increases energy expenditure and reduces body weight gain
title_fullStr Hypothalamic over-expression of VGF in the Siberian hamster increases energy expenditure and reduces body weight gain
title_full_unstemmed Hypothalamic over-expression of VGF in the Siberian hamster increases energy expenditure and reduces body weight gain
title_short Hypothalamic over-expression of VGF in the Siberian hamster increases energy expenditure and reduces body weight gain
title_sort hypothalamic over-expression of vgf in the siberian hamster increases energy expenditure and reduces body weight gain
topic Siberian hamsters
VGF
hypothalamus
adeno-associated viral virus
url https://eprints.nottingham.ac.uk/40849/
https://eprints.nottingham.ac.uk/40849/
https://eprints.nottingham.ac.uk/40849/