Dormant and after-ripened Arabidopsis thaliana seeds are distinguished by early transcriptional differences in the imbibed state

Seed dormancy is a genetically controlled block preventing the germination of imbibed seeds in favorable conditions. It requires a period of dry storage (after-ripening) or certain environmental conditions to be overcome. Dormancy is an important seed trait, which is under selective pressure, to con...

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Main Authors: Dekkers, Bas J.W., Pearce, Simon P., van Bolderen-Veldkamp, R.P.M., Holdsworth, Michael J., Bentsink, Leónie
Format: Article
Published: Frontiers Media 2016
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Online Access:https://eprints.nottingham.ac.uk/38963/
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author Dekkers, Bas J.W.
Pearce, Simon P.
van Bolderen-Veldkamp, R.P.M.
Holdsworth, Michael J.
Bentsink, Leónie
author_facet Dekkers, Bas J.W.
Pearce, Simon P.
van Bolderen-Veldkamp, R.P.M.
Holdsworth, Michael J.
Bentsink, Leónie
author_sort Dekkers, Bas J.W.
building Nottingham Research Data Repository
collection Online Access
description Seed dormancy is a genetically controlled block preventing the germination of imbibed seeds in favorable conditions. It requires a period of dry storage (after-ripening) or certain environmental conditions to be overcome. Dormancy is an important seed trait, which is under selective pressure, to control the seasonal timing of seed germination. Dormant and non-dormant (after-ripened) seeds are characterized by large sets of differentially expressed genes. However, little information is available concerning the temporal and spatial transcriptional changes during early stages of rehydration in dormant and nondormant seeds. We employed genome-wide transcriptome analysis on seeds of the model plant Arabidopsis thaliana to investigate transcriptional changes in dry seeds upon rehydration. We analyzed gene expression of dormant and after-ripened seeds of the Cvi accession over four time points and two seed compartments (the embryo and surrounding single cell layer endosperm), during the first 24 h after sowing. This work provides a global view of gene expression changes in dormant and non-dormant seeds with temporal and spatial detail, and these may be visualized via a web accessible tool (http://www.wageningenseedlab.nl/resources). A large proportion of transcripts change similarly in both dormant and non-dormant seeds upon rehydration, however, the first differences in transcript abundances become visible shortly after the initiation of imbibition, indicating that changes induced by after-ripening are detected and responded to rapidly upon rehydration. We identified several gene expression profiles which contribute to differential gene expression between dormant and non-dormant samples. Genes with enhanced expression in the endosperm of dormant seeds were overrepresented for stress-related Gene Ontology categories, suggesting a protective role for the endosperm against biotic and abiotic stress to support persistence of the dormant seed in its environment.
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spelling nottingham-389632020-05-04T18:05:18Z https://eprints.nottingham.ac.uk/38963/ Dormant and after-ripened Arabidopsis thaliana seeds are distinguished by early transcriptional differences in the imbibed state Dekkers, Bas J.W. Pearce, Simon P. van Bolderen-Veldkamp, R.P.M. Holdsworth, Michael J. Bentsink, Leónie Seed dormancy is a genetically controlled block preventing the germination of imbibed seeds in favorable conditions. It requires a period of dry storage (after-ripening) or certain environmental conditions to be overcome. Dormancy is an important seed trait, which is under selective pressure, to control the seasonal timing of seed germination. Dormant and non-dormant (after-ripened) seeds are characterized by large sets of differentially expressed genes. However, little information is available concerning the temporal and spatial transcriptional changes during early stages of rehydration in dormant and nondormant seeds. We employed genome-wide transcriptome analysis on seeds of the model plant Arabidopsis thaliana to investigate transcriptional changes in dry seeds upon rehydration. We analyzed gene expression of dormant and after-ripened seeds of the Cvi accession over four time points and two seed compartments (the embryo and surrounding single cell layer endosperm), during the first 24 h after sowing. This work provides a global view of gene expression changes in dormant and non-dormant seeds with temporal and spatial detail, and these may be visualized via a web accessible tool (http://www.wageningenseedlab.nl/resources). A large proportion of transcripts change similarly in both dormant and non-dormant seeds upon rehydration, however, the first differences in transcript abundances become visible shortly after the initiation of imbibition, indicating that changes induced by after-ripening are detected and responded to rapidly upon rehydration. We identified several gene expression profiles which contribute to differential gene expression between dormant and non-dormant samples. Genes with enhanced expression in the endosperm of dormant seeds were overrepresented for stress-related Gene Ontology categories, suggesting a protective role for the endosperm against biotic and abiotic stress to support persistence of the dormant seed in its environment. Frontiers Media 2016-08-30 Article PeerReviewed Dekkers, Bas J.W., Pearce, Simon P., van Bolderen-Veldkamp, R.P.M., Holdsworth, Michael J. and Bentsink, Leónie (2016) Dormant and after-ripened Arabidopsis thaliana seeds are distinguished by early transcriptional differences in the imbibed state. Frontiers in Plant Science, 7 . 1323/1-1323/15. ISSN 1664-462X After-ripening Arabidopsis Dormancy Embryo Endosperm Seed Transcriptome http://dx.doi.org/10.3389/fpls.2016.01323 doi:10.3389/fpls.2016.01323 doi:10.3389/fpls.2016.01323
spellingShingle After-ripening
Arabidopsis
Dormancy
Embryo
Endosperm
Seed
Transcriptome
Dekkers, Bas J.W.
Pearce, Simon P.
van Bolderen-Veldkamp, R.P.M.
Holdsworth, Michael J.
Bentsink, Leónie
Dormant and after-ripened Arabidopsis thaliana seeds are distinguished by early transcriptional differences in the imbibed state
title Dormant and after-ripened Arabidopsis thaliana seeds are distinguished by early transcriptional differences in the imbibed state
title_full Dormant and after-ripened Arabidopsis thaliana seeds are distinguished by early transcriptional differences in the imbibed state
title_fullStr Dormant and after-ripened Arabidopsis thaliana seeds are distinguished by early transcriptional differences in the imbibed state
title_full_unstemmed Dormant and after-ripened Arabidopsis thaliana seeds are distinguished by early transcriptional differences in the imbibed state
title_short Dormant and after-ripened Arabidopsis thaliana seeds are distinguished by early transcriptional differences in the imbibed state
title_sort dormant and after-ripened arabidopsis thaliana seeds are distinguished by early transcriptional differences in the imbibed state
topic After-ripening
Arabidopsis
Dormancy
Embryo
Endosperm
Seed
Transcriptome
url https://eprints.nottingham.ac.uk/38963/
https://eprints.nottingham.ac.uk/38963/
https://eprints.nottingham.ac.uk/38963/