Biphasic regulation of the transcription factor ABORTED MICROSPORES (AMS) is essential for tapetum and pollen development in Arabidopsis

Viable pollen is essential for plant reproduction and crop yield. Its production requires coordinated expression at specific stages during anther development, involving early meiosis-associated events and late pollen wall formation. The ABORTED MICROSPORE (AMS) transcription factor is a master regul...

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Main Authors: Ferguson, Alison, Pearce, Simon, Band, Leah R., Yang, Caiyun, Ferjentsikova, Ivana, King, John, Yuan, Zheng, Zhang, Dabing, Wilson, Zoe A.
Format: Article
Published: Wiley 2016
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Online Access:https://eprints.nottingham.ac.uk/37902/
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author Ferguson, Alison
Pearce, Simon
Band, Leah R.
Yang, Caiyun
Ferjentsikova, Ivana
King, John
Yuan, Zheng
Zhang, Dabing
Wilson, Zoe A.
author_facet Ferguson, Alison
Pearce, Simon
Band, Leah R.
Yang, Caiyun
Ferjentsikova, Ivana
King, John
Yuan, Zheng
Zhang, Dabing
Wilson, Zoe A.
author_sort Ferguson, Alison
building Nottingham Research Data Repository
collection Online Access
description Viable pollen is essential for plant reproduction and crop yield. Its production requires coordinated expression at specific stages during anther development, involving early meiosis-associated events and late pollen wall formation. The ABORTED MICROSPORE (AMS) transcription factor is a master regulator of sporopollenin biosynthesis, secretion and pollen wall formation in Arabidopsis. Here we show it has complex regulation and additional essential roles earlier in pollen formation. • An inducible-AMS reporter was created for functional rescue, protein expression pattern analysis and to distinguish between direct and indirect targets. Mathematical modelling was used to create regulatory networks based on wildtype RNA and protein expression. • Dual activity of AMS was defined by biphasic protein expression in anther tapetal cells, with an initial peak around pollen meiosis and then later during pollen wall development. Direct AMS-regulated targets exhibit temporal regulation, indicating additional factors are associated with their regulation. • We demonstrate that AMS biphasic expression is essential for pollen development and defines distinct functional activities during early and late pollen development. Mathematical modelling suggests AMS may competitively form a protein complex with other tapetum-expressed transcription factors, and that biphasic regulation is due to repression of upstream regulators and promotion of AMS protein degradation.
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spelling nottingham-379022020-05-04T18:15:11Z https://eprints.nottingham.ac.uk/37902/ Biphasic regulation of the transcription factor ABORTED MICROSPORES (AMS) is essential for tapetum and pollen development in Arabidopsis Ferguson, Alison Pearce, Simon Band, Leah R. Yang, Caiyun Ferjentsikova, Ivana King, John Yuan, Zheng Zhang, Dabing Wilson, Zoe A. Viable pollen is essential for plant reproduction and crop yield. Its production requires coordinated expression at specific stages during anther development, involving early meiosis-associated events and late pollen wall formation. The ABORTED MICROSPORE (AMS) transcription factor is a master regulator of sporopollenin biosynthesis, secretion and pollen wall formation in Arabidopsis. Here we show it has complex regulation and additional essential roles earlier in pollen formation. • An inducible-AMS reporter was created for functional rescue, protein expression pattern analysis and to distinguish between direct and indirect targets. Mathematical modelling was used to create regulatory networks based on wildtype RNA and protein expression. • Dual activity of AMS was defined by biphasic protein expression in anther tapetal cells, with an initial peak around pollen meiosis and then later during pollen wall development. Direct AMS-regulated targets exhibit temporal regulation, indicating additional factors are associated with their regulation. • We demonstrate that AMS biphasic expression is essential for pollen development and defines distinct functional activities during early and late pollen development. Mathematical modelling suggests AMS may competitively form a protein complex with other tapetum-expressed transcription factors, and that biphasic regulation is due to repression of upstream regulators and promotion of AMS protein degradation. Wiley 2016-10-27 Article PeerReviewed Ferguson, Alison, Pearce, Simon, Band, Leah R., Yang, Caiyun, Ferjentsikova, Ivana, King, John, Yuan, Zheng, Zhang, Dabing and Wilson, Zoe A. (2016) Biphasic regulation of the transcription factor ABORTED MICROSPORES (AMS) is essential for tapetum and pollen development in Arabidopsis. New Phytologist, 213 . pp. 778-790. ISSN 1469-8137 Aborted microspore (AMS) anther development Arabidopsis thaliana pollen development regulatory network modelling tapetum http://onlinelibrary.wiley.com/doi/10.1111/nph.14200/abstract doi:10.1111/nph.14200 doi:10.1111/nph.14200
spellingShingle Aborted microspore (AMS)
anther development
Arabidopsis thaliana
pollen development
regulatory network modelling
tapetum
Ferguson, Alison
Pearce, Simon
Band, Leah R.
Yang, Caiyun
Ferjentsikova, Ivana
King, John
Yuan, Zheng
Zhang, Dabing
Wilson, Zoe A.
Biphasic regulation of the transcription factor ABORTED MICROSPORES (AMS) is essential for tapetum and pollen development in Arabidopsis
title Biphasic regulation of the transcription factor ABORTED MICROSPORES (AMS) is essential for tapetum and pollen development in Arabidopsis
title_full Biphasic regulation of the transcription factor ABORTED MICROSPORES (AMS) is essential for tapetum and pollen development in Arabidopsis
title_fullStr Biphasic regulation of the transcription factor ABORTED MICROSPORES (AMS) is essential for tapetum and pollen development in Arabidopsis
title_full_unstemmed Biphasic regulation of the transcription factor ABORTED MICROSPORES (AMS) is essential for tapetum and pollen development in Arabidopsis
title_short Biphasic regulation of the transcription factor ABORTED MICROSPORES (AMS) is essential for tapetum and pollen development in Arabidopsis
title_sort biphasic regulation of the transcription factor aborted microspores (ams) is essential for tapetum and pollen development in arabidopsis
topic Aborted microspore (AMS)
anther development
Arabidopsis thaliana
pollen development
regulatory network modelling
tapetum
url https://eprints.nottingham.ac.uk/37902/
https://eprints.nottingham.ac.uk/37902/
https://eprints.nottingham.ac.uk/37902/