TP53 copy number expansion is associated with the evolution of increased body size and an enhanced DNA damage response in elephants.

A major constraint on the evolution of large body sizes in animals is an increased risk of developing cancer. There is no correlation, however, between body size and cancer risk. This lack of correlation is often referred to as 'Peto's Paradox'. Here, we show that the elephant genome...

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Main Authors: Sulak, Michael, Fong, Lindsey, Mika, Katelyn, Chigurupati, Sravanthi, Yon, Lisa, Mongan, Nigel P., Emes, Richard D., Lynch, Vincent J.
Format: Article
Published: eLife Sciences Publications 2016
Online Access:https://eprints.nottingham.ac.uk/37731/
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author Sulak, Michael
Fong, Lindsey
Mika, Katelyn
Chigurupati, Sravanthi
Yon, Lisa
Mongan, Nigel P.
Emes, Richard D.
Lynch, Vincent J.
author_facet Sulak, Michael
Fong, Lindsey
Mika, Katelyn
Chigurupati, Sravanthi
Yon, Lisa
Mongan, Nigel P.
Emes, Richard D.
Lynch, Vincent J.
author_sort Sulak, Michael
building Nottingham Research Data Repository
collection Online Access
description A major constraint on the evolution of large body sizes in animals is an increased risk of developing cancer. There is no correlation, however, between body size and cancer risk. This lack of correlation is often referred to as 'Peto's Paradox'. Here, we show that the elephant genome encodes 20 copies of the tumor suppressor gene TP53 and that the increase in TP53 copy number occurred coincident with the evolution of large body sizes, the evolution of extreme sensitivity to genotoxic stress, and a hyperactive TP53 signaling pathway in the elephant (Proboscidean) lineage. Furthermore, we show that several of the TP53 retrogenes (TP53RTGs) are transcribed and likely translated. While TP53RTGs do not appear to directly function as transcription factors, they do contribute to the enhanced sensitivity of elephant cells to DNA damage and the induction of apoptosis by regulating activity of the TP53 signaling pathway. These results suggest that an increase in the copy number of TP53 may have played a direct role in the evolution of very large body sizes and the resolution of Peto's paradox in Proboscideans.
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spelling nottingham-377312020-05-04T18:11:45Z https://eprints.nottingham.ac.uk/37731/ TP53 copy number expansion is associated with the evolution of increased body size and an enhanced DNA damage response in elephants. Sulak, Michael Fong, Lindsey Mika, Katelyn Chigurupati, Sravanthi Yon, Lisa Mongan, Nigel P. Emes, Richard D. Lynch, Vincent J. A major constraint on the evolution of large body sizes in animals is an increased risk of developing cancer. There is no correlation, however, between body size and cancer risk. This lack of correlation is often referred to as 'Peto's Paradox'. Here, we show that the elephant genome encodes 20 copies of the tumor suppressor gene TP53 and that the increase in TP53 copy number occurred coincident with the evolution of large body sizes, the evolution of extreme sensitivity to genotoxic stress, and a hyperactive TP53 signaling pathway in the elephant (Proboscidean) lineage. Furthermore, we show that several of the TP53 retrogenes (TP53RTGs) are transcribed and likely translated. While TP53RTGs do not appear to directly function as transcription factors, they do contribute to the enhanced sensitivity of elephant cells to DNA damage and the induction of apoptosis by regulating activity of the TP53 signaling pathway. These results suggest that an increase in the copy number of TP53 may have played a direct role in the evolution of very large body sizes and the resolution of Peto's paradox in Proboscideans. eLife Sciences Publications 2016-09-19 Article PeerReviewed Sulak, Michael, Fong, Lindsey, Mika, Katelyn, Chigurupati, Sravanthi, Yon, Lisa, Mongan, Nigel P., Emes, Richard D. and Lynch, Vincent J. (2016) TP53 copy number expansion is associated with the evolution of increased body size and an enhanced DNA damage response in elephants. eLife, 5 (e11994). ISSN 2050-084X https://elifesciences.org/content/5/e11994 doi:10.7554/eLife.11994 doi:10.7554/eLife.11994
spellingShingle Sulak, Michael
Fong, Lindsey
Mika, Katelyn
Chigurupati, Sravanthi
Yon, Lisa
Mongan, Nigel P.
Emes, Richard D.
Lynch, Vincent J.
TP53 copy number expansion is associated with the evolution of increased body size and an enhanced DNA damage response in elephants.
title TP53 copy number expansion is associated with the evolution of increased body size and an enhanced DNA damage response in elephants.
title_full TP53 copy number expansion is associated with the evolution of increased body size and an enhanced DNA damage response in elephants.
title_fullStr TP53 copy number expansion is associated with the evolution of increased body size and an enhanced DNA damage response in elephants.
title_full_unstemmed TP53 copy number expansion is associated with the evolution of increased body size and an enhanced DNA damage response in elephants.
title_short TP53 copy number expansion is associated with the evolution of increased body size and an enhanced DNA damage response in elephants.
title_sort tp53 copy number expansion is associated with the evolution of increased body size and an enhanced dna damage response in elephants.
url https://eprints.nottingham.ac.uk/37731/
https://eprints.nottingham.ac.uk/37731/
https://eprints.nottingham.ac.uk/37731/