Primordial germ cells: the first cell lineage or the last cells standing?

Embryos of many animal models express germ line determinants that suppress transcription and mediate early germ line commitment, which occurs before the somatic cell lineages are established. However, not all animals segregate their germ line in this manner. The ‘last cell standing’ model describes...

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Main Authors: Johnson, Andrew D., Alberio, Ramiro
Format: Article
Published: Company of Biologists 2015
Subjects:
Online Access:https://eprints.nottingham.ac.uk/37066/
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author Johnson, Andrew D.
Alberio, Ramiro
author_facet Johnson, Andrew D.
Alberio, Ramiro
author_sort Johnson, Andrew D.
building Nottingham Research Data Repository
collection Online Access
description Embryos of many animal models express germ line determinants that suppress transcription and mediate early germ line commitment, which occurs before the somatic cell lineages are established. However, not all animals segregate their germ line in this manner. The ‘last cell standing’ model describes primordial germ cell (PGC) development in axolotls, in which PGCs are maintained by an extracellular signalling niche, and germ line commitment occurs after gastrulation. Here, we propose that this ‘stochastic’ mode of PGC specification is conserved in vertebrates, including non-rodent mammals. We postulate that early germ line segregation liberates genetic regulatory networks for somatic development to evolve, and that it therefore emerged repeatedly in the animal kingdom in response to natural selection.
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spelling nottingham-370662020-05-04T17:14:53Z https://eprints.nottingham.ac.uk/37066/ Primordial germ cells: the first cell lineage or the last cells standing? Johnson, Andrew D. Alberio, Ramiro Embryos of many animal models express germ line determinants that suppress transcription and mediate early germ line commitment, which occurs before the somatic cell lineages are established. However, not all animals segregate their germ line in this manner. The ‘last cell standing’ model describes primordial germ cell (PGC) development in axolotls, in which PGCs are maintained by an extracellular signalling niche, and germ line commitment occurs after gastrulation. Here, we propose that this ‘stochastic’ mode of PGC specification is conserved in vertebrates, including non-rodent mammals. We postulate that early germ line segregation liberates genetic regulatory networks for somatic development to evolve, and that it therefore emerged repeatedly in the animal kingdom in response to natural selection. Company of Biologists 2015-08-18 Article PeerReviewed Johnson, Andrew D. and Alberio, Ramiro (2015) Primordial germ cells: the first cell lineage or the last cells standing? Development, 142 . pp. 2730-2739. ISSN 1477-9129 Amphibian embryo Evolvability Germ plasm Mammalian embryo Pluripotency Primordial germ cell PGC http://dev.biologists.org/content/142/16/2730 doi: 10.1242/dev.113993 doi: 10.1242/dev.113993
spellingShingle Amphibian embryo
Evolvability
Germ plasm
Mammalian embryo
Pluripotency
Primordial germ cell
PGC
Johnson, Andrew D.
Alberio, Ramiro
Primordial germ cells: the first cell lineage or the last cells standing?
title Primordial germ cells: the first cell lineage or the last cells standing?
title_full Primordial germ cells: the first cell lineage or the last cells standing?
title_fullStr Primordial germ cells: the first cell lineage or the last cells standing?
title_full_unstemmed Primordial germ cells: the first cell lineage or the last cells standing?
title_short Primordial germ cells: the first cell lineage or the last cells standing?
title_sort primordial germ cells: the first cell lineage or the last cells standing?
topic Amphibian embryo
Evolvability
Germ plasm
Mammalian embryo
Pluripotency
Primordial germ cell
PGC
url https://eprints.nottingham.ac.uk/37066/
https://eprints.nottingham.ac.uk/37066/
https://eprints.nottingham.ac.uk/37066/