Functional characterisation of small signalling peptides in the regulation of Arabidopsis thaliana development

Root architecture governs the uptake of water and nutrients from the soil and thus is essential for plant growth and survival. The control of lateral root branching is a crucial aspect of determining root architecture, and is a process largely controlled by the phytohormone auxin, which promotes lat...

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Main Author: Smith, Stephanie
Format: Thesis (University of Nottingham only)
Language:English
Published: 2016
Subjects:
Online Access:https://eprints.nottingham.ac.uk/33853/
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author Smith, Stephanie
author_facet Smith, Stephanie
author_sort Smith, Stephanie
building Nottingham Research Data Repository
collection Online Access
description Root architecture governs the uptake of water and nutrients from the soil and thus is essential for plant growth and survival. The control of lateral root branching is a crucial aspect of determining root architecture, and is a process largely controlled by the phytohormone auxin, which promotes lateral root organogenesis from founder cells within the vasculature and also subsequent lateral root elongation. A small posttranslationally modified peptide, CTERMINALLY ENCODED PEPTIDE1 (CEP1) is a bioactive 15-amino acid peptide cleaved from a larger precursor protein previously shown to negatively affect primary root elongation and lateral root branching in Arabidopsis thaliana. Within this thesis, genetic, biochemical and pharmacological approaches are used to demonstrate that another member of the CEP family, CEP5, reduces primary root length and negatively regulates lateral root branching in Arabidopsis thaliana with minor effects on aboveground architecture. CEP5 is also demonstrated to have an antagonistic relationship with auxin, with increased abundance of CEP5 leading to downstream stabilisation of the auxin signalling repressor AUXIN/INDOLE ACETIC ACID (AUX/IAA) proteins. Correspondingly, reduced abundance of CEP5 increases auxin responses, including enhanced lateral root progression. These studies suggest CEP5 regulates lateral root branching through attenuation of auxin responses.
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spelling nottingham-338532025-02-28T13:29:55Z https://eprints.nottingham.ac.uk/33853/ Functional characterisation of small signalling peptides in the regulation of Arabidopsis thaliana development Smith, Stephanie Root architecture governs the uptake of water and nutrients from the soil and thus is essential for plant growth and survival. The control of lateral root branching is a crucial aspect of determining root architecture, and is a process largely controlled by the phytohormone auxin, which promotes lateral root organogenesis from founder cells within the vasculature and also subsequent lateral root elongation. A small posttranslationally modified peptide, CTERMINALLY ENCODED PEPTIDE1 (CEP1) is a bioactive 15-amino acid peptide cleaved from a larger precursor protein previously shown to negatively affect primary root elongation and lateral root branching in Arabidopsis thaliana. Within this thesis, genetic, biochemical and pharmacological approaches are used to demonstrate that another member of the CEP family, CEP5, reduces primary root length and negatively regulates lateral root branching in Arabidopsis thaliana with minor effects on aboveground architecture. CEP5 is also demonstrated to have an antagonistic relationship with auxin, with increased abundance of CEP5 leading to downstream stabilisation of the auxin signalling repressor AUXIN/INDOLE ACETIC ACID (AUX/IAA) proteins. Correspondingly, reduced abundance of CEP5 increases auxin responses, including enhanced lateral root progression. These studies suggest CEP5 regulates lateral root branching through attenuation of auxin responses. 2016-07-21 Thesis (University of Nottingham only) NonPeerReviewed application/pdf en arr https://eprints.nottingham.ac.uk/33853/1/StephanieSmithThesisCorrected1.pdf Smith, Stephanie (2016) Functional characterisation of small signalling peptides in the regulation of Arabidopsis thaliana development. PhD thesis, University of Nottingham. CEP Arabidopsis thaliana lateral root root architecture root development nitrate sensing auxin small postranslationally modified peptides
spellingShingle CEP
Arabidopsis thaliana
lateral root
root architecture
root development
nitrate sensing
auxin
small postranslationally modified peptides
Smith, Stephanie
Functional characterisation of small signalling peptides in the regulation of Arabidopsis thaliana development
title Functional characterisation of small signalling peptides in the regulation of Arabidopsis thaliana development
title_full Functional characterisation of small signalling peptides in the regulation of Arabidopsis thaliana development
title_fullStr Functional characterisation of small signalling peptides in the regulation of Arabidopsis thaliana development
title_full_unstemmed Functional characterisation of small signalling peptides in the regulation of Arabidopsis thaliana development
title_short Functional characterisation of small signalling peptides in the regulation of Arabidopsis thaliana development
title_sort functional characterisation of small signalling peptides in the regulation of arabidopsis thaliana development
topic CEP
Arabidopsis thaliana
lateral root
root architecture
root development
nitrate sensing
auxin
small postranslationally modified peptides
url https://eprints.nottingham.ac.uk/33853/