Synergism between cAMP and PPARγ signalling in the initiation of UCP1 gene expression in HIB1B brown adipocytes

Expression of the brown adipocyte-specific gene, uncoupling protein 1 (UCP1), is increased by both PPAR stimulation and cAMP activation through their ability to stimulate the expression of the PPAR coactivator PGC1. In HIB1B brown preadipocytes, combination of the PPAR agonist, rosiglitazone, and th...

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Main Authors: Chen, H.Y., Liu, Q., Salter, Andrew M., Lomax, Michael A.
Format: Article
Published: Hindawi Publishing Corporation 2013
Online Access:https://eprints.nottingham.ac.uk/3170/
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author Chen, H.Y.
Liu, Q.
Salter, Andrew M.
Lomax, Michael A.
author_facet Chen, H.Y.
Liu, Q.
Salter, Andrew M.
Lomax, Michael A.
author_sort Chen, H.Y.
building Nottingham Research Data Repository
collection Online Access
description Expression of the brown adipocyte-specific gene, uncoupling protein 1 (UCP1), is increased by both PPAR stimulation and cAMP activation through their ability to stimulate the expression of the PPAR coactivator PGC1. In HIB1B brown preadipocytes, combination of the PPAR agonist, rosiglitazone, and the cAMP stimulator forskolin synergistically increased UCP1 mRNA expression, but PGC1 expression was only increased additively by the two drugs. The PPAR antagonist, GW9662, and the PKA inhibitor, H89, both inhibited UCP1 expression stimulated by rosiglitazone and forskolin but PGC1 expression was not altered to the same extent. Reporter studies demonstrated that combined rosiglitazone and forskolin synergistically activated transcription from a full length 3.1 kbp UCP1 luciferase promoter construct, but the response was only additive and much reduced when a minimal 260 bp proximal UCP1 promoter was examined. Rosiglitazone and forskolin in combination were able to synergistically stimulate promoters comprising of tandem repeats of either PPREs or CREs. We conclude that rosiglitazone and forskolin act together to synergistically activate the UCP1 promoter directly rather than by increasing PGC1 expression and by a mechanism involving cross-talk between the signalling systems regulating the CRE and PPRE on the promoters.
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spelling nottingham-31702020-05-04T20:19:55Z https://eprints.nottingham.ac.uk/3170/ Synergism between cAMP and PPARγ signalling in the initiation of UCP1 gene expression in HIB1B brown adipocytes Chen, H.Y. Liu, Q. Salter, Andrew M. Lomax, Michael A. Expression of the brown adipocyte-specific gene, uncoupling protein 1 (UCP1), is increased by both PPAR stimulation and cAMP activation through their ability to stimulate the expression of the PPAR coactivator PGC1. In HIB1B brown preadipocytes, combination of the PPAR agonist, rosiglitazone, and the cAMP stimulator forskolin synergistically increased UCP1 mRNA expression, but PGC1 expression was only increased additively by the two drugs. The PPAR antagonist, GW9662, and the PKA inhibitor, H89, both inhibited UCP1 expression stimulated by rosiglitazone and forskolin but PGC1 expression was not altered to the same extent. Reporter studies demonstrated that combined rosiglitazone and forskolin synergistically activated transcription from a full length 3.1 kbp UCP1 luciferase promoter construct, but the response was only additive and much reduced when a minimal 260 bp proximal UCP1 promoter was examined. Rosiglitazone and forskolin in combination were able to synergistically stimulate promoters comprising of tandem repeats of either PPREs or CREs. We conclude that rosiglitazone and forskolin act together to synergistically activate the UCP1 promoter directly rather than by increasing PGC1 expression and by a mechanism involving cross-talk between the signalling systems regulating the CRE and PPRE on the promoters. Hindawi Publishing Corporation 2013 Article PeerReviewed Chen, H.Y., Liu, Q., Salter, Andrew M. and Lomax, Michael A. (2013) Synergism between cAMP and PPARγ signalling in the initiation of UCP1 gene expression in HIB1B brown adipocytes. PPAR Research, 2013 (476049). ISSN 1687-4757 http://www.hindawi.com/journals/ppar/2013/476049/ doi:10.1155/2013/476049 doi:10.1155/2013/476049
spellingShingle Chen, H.Y.
Liu, Q.
Salter, Andrew M.
Lomax, Michael A.
Synergism between cAMP and PPARγ signalling in the initiation of UCP1 gene expression in HIB1B brown adipocytes
title Synergism between cAMP and PPARγ signalling in the initiation of UCP1 gene expression in HIB1B brown adipocytes
title_full Synergism between cAMP and PPARγ signalling in the initiation of UCP1 gene expression in HIB1B brown adipocytes
title_fullStr Synergism between cAMP and PPARγ signalling in the initiation of UCP1 gene expression in HIB1B brown adipocytes
title_full_unstemmed Synergism between cAMP and PPARγ signalling in the initiation of UCP1 gene expression in HIB1B brown adipocytes
title_short Synergism between cAMP and PPARγ signalling in the initiation of UCP1 gene expression in HIB1B brown adipocytes
title_sort synergism between camp and pparγ signalling in the initiation of ucp1 gene expression in hib1b brown adipocytes
url https://eprints.nottingham.ac.uk/3170/
https://eprints.nottingham.ac.uk/3170/
https://eprints.nottingham.ac.uk/3170/