Asymmetric competition causes multimodal size distributions in spatially structured populations

Plant sizes within populations often exhibit multimodal distributions, even when all individuals are the same age and have experienced identical conditions. To establish the causes of this, we created an individual-based model simulating the growth of trees in a spatially explicit framework, which w...

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Main Authors: Velázquez, Jorge, Allen, Robert B., Coomes, David A., Eichhorn, Markus P.
Format: Article
Published: Royal Society 2016
Subjects:
Online Access:https://eprints.nottingham.ac.uk/31516/
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author Velázquez, Jorge
Allen, Robert B.
Coomes, David A.
Eichhorn, Markus P.
author_facet Velázquez, Jorge
Allen, Robert B.
Coomes, David A.
Eichhorn, Markus P.
author_sort Velázquez, Jorge
building Nottingham Research Data Repository
collection Online Access
description Plant sizes within populations often exhibit multimodal distributions, even when all individuals are the same age and have experienced identical conditions. To establish the causes of this, we created an individual-based model simulating the growth of trees in a spatially explicit framework, which was parametrized using data from a long-term study of forest stands in New Zealand. First, we demonstrate that asymmetric resource competition is a necessary condition for the formation of multimodal size distributions within cohorts. By contrast, the legacy of small-scale clustering during recruitment is transient and quickly overwhelmed by density-dependent mortality. Complex multi-layered size distributions are generated when established individuals are restricted in the spatial domain within which they can capture resources. The number of modes reveals the effective number of direct competitors, while the separation and spread of modes are influenced by distances among established individuals. Asymmetric competition within local neighbourhoods can therefore generate a range of complex size distributions within even-aged cohorts.
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spelling nottingham-315162020-05-04T17:29:57Z https://eprints.nottingham.ac.uk/31516/ Asymmetric competition causes multimodal size distributions in spatially structured populations Velázquez, Jorge Allen, Robert B. Coomes, David A. Eichhorn, Markus P. Plant sizes within populations often exhibit multimodal distributions, even when all individuals are the same age and have experienced identical conditions. To establish the causes of this, we created an individual-based model simulating the growth of trees in a spatially explicit framework, which was parametrized using data from a long-term study of forest stands in New Zealand. First, we demonstrate that asymmetric resource competition is a necessary condition for the formation of multimodal size distributions within cohorts. By contrast, the legacy of small-scale clustering during recruitment is transient and quickly overwhelmed by density-dependent mortality. Complex multi-layered size distributions are generated when established individuals are restricted in the spatial domain within which they can capture resources. The number of modes reveals the effective number of direct competitors, while the separation and spread of modes are influenced by distances among established individuals. Asymmetric competition within local neighbourhoods can therefore generate a range of complex size distributions within even-aged cohorts. Royal Society 2016-01-27 Article PeerReviewed Velázquez, Jorge, Allen, Robert B., Coomes, David A. and Eichhorn, Markus P. (2016) Asymmetric competition causes multimodal size distributions in spatially structured populations. Proceedings of the Royal Society B: Biological Sciences, 283 (1823). ISSN 1471-2954 bimodality individual-based model forests Fuscospora cliffortioides southern beech zone-of-influence http://rspb.royalsocietypublishing.org/content/283/1823/20152404 doi:10.1098/rspb.2015.2404 doi:10.1098/rspb.2015.2404
spellingShingle bimodality
individual-based model
forests
Fuscospora cliffortioides
southern beech
zone-of-influence
Velázquez, Jorge
Allen, Robert B.
Coomes, David A.
Eichhorn, Markus P.
Asymmetric competition causes multimodal size distributions in spatially structured populations
title Asymmetric competition causes multimodal size distributions in spatially structured populations
title_full Asymmetric competition causes multimodal size distributions in spatially structured populations
title_fullStr Asymmetric competition causes multimodal size distributions in spatially structured populations
title_full_unstemmed Asymmetric competition causes multimodal size distributions in spatially structured populations
title_short Asymmetric competition causes multimodal size distributions in spatially structured populations
title_sort asymmetric competition causes multimodal size distributions in spatially structured populations
topic bimodality
individual-based model
forests
Fuscospora cliffortioides
southern beech
zone-of-influence
url https://eprints.nottingham.ac.uk/31516/
https://eprints.nottingham.ac.uk/31516/
https://eprints.nottingham.ac.uk/31516/