Cooperation, quorum sensing, and evolution of virulence in Staphylococcus aureus

The virulence and fitness in vivo of the major human pathogen Staphylococcus aureus are associated with a cell-to-cell signaling mechanism known as quorum sensing (QS). QS coordinates the production of virulence factors via the production and sensing of autoinducing peptide (AIP) signal molecules by...

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Main Authors: Pollitt, Eric J.G., West, Stuart A., Crusz, Shanika A., Burton-Chellew, Maxwell N., Diggle, Stephen P.
Format: Article
Published: ASM Press 2014
Online Access:https://eprints.nottingham.ac.uk/2991/
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author Pollitt, Eric J.G.
West, Stuart A.
Crusz, Shanika A.
Burton-Chellew, Maxwell N.
Diggle, Stephen P.
author_facet Pollitt, Eric J.G.
West, Stuart A.
Crusz, Shanika A.
Burton-Chellew, Maxwell N.
Diggle, Stephen P.
author_sort Pollitt, Eric J.G.
building Nottingham Research Data Repository
collection Online Access
description The virulence and fitness in vivo of the major human pathogen Staphylococcus aureus are associated with a cell-to-cell signaling mechanism known as quorum sensing (QS). QS coordinates the production of virulence factors via the production and sensing of autoinducing peptide (AIP) signal molecules by the agr locus. Here we show, in a wax moth larva virulence model, that (i) QS in S. aureus is a cooperative social trait that provides a benefit to the local population of cells, (ii) agr mutants, which do not pro- duce or respond to QS signal, are able to exploit the benefits provided by the QS of others (“cheat”), allowing them to increase in frequency when in mixed populations with cooperators, (iii) these social interactions between cells determine virulence, with the host mortality rate being negatively correlated to the percentage of agr mutants (“cheats”) in a population, and (iv) a higher within-host relatedness (lower strain diversity) selects for QS and hence higher virulence. Our results provide an explanation for why agr mutants show reduced virulence in animal models but can be isolated from infections of humans. More generally, by providing the first evidence that QS is a cooperative social behavior in a Gram-positive bacterium, our results suggest conver- gent, and potentially widespread, evolution for signaling to coordinate cooperation in bacteria.
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spelling nottingham-29912020-05-04T20:17:36Z https://eprints.nottingham.ac.uk/2991/ Cooperation, quorum sensing, and evolution of virulence in Staphylococcus aureus Pollitt, Eric J.G. West, Stuart A. Crusz, Shanika A. Burton-Chellew, Maxwell N. Diggle, Stephen P. The virulence and fitness in vivo of the major human pathogen Staphylococcus aureus are associated with a cell-to-cell signaling mechanism known as quorum sensing (QS). QS coordinates the production of virulence factors via the production and sensing of autoinducing peptide (AIP) signal molecules by the agr locus. Here we show, in a wax moth larva virulence model, that (i) QS in S. aureus is a cooperative social trait that provides a benefit to the local population of cells, (ii) agr mutants, which do not pro- duce or respond to QS signal, are able to exploit the benefits provided by the QS of others (“cheat”), allowing them to increase in frequency when in mixed populations with cooperators, (iii) these social interactions between cells determine virulence, with the host mortality rate being negatively correlated to the percentage of agr mutants (“cheats”) in a population, and (iv) a higher within-host relatedness (lower strain diversity) selects for QS and hence higher virulence. Our results provide an explanation for why agr mutants show reduced virulence in animal models but can be isolated from infections of humans. More generally, by providing the first evidence that QS is a cooperative social behavior in a Gram-positive bacterium, our results suggest conver- gent, and potentially widespread, evolution for signaling to coordinate cooperation in bacteria. ASM Press 2014 Article PeerReviewed Pollitt, Eric J.G., West, Stuart A., Crusz, Shanika A., Burton-Chellew, Maxwell N. and Diggle, Stephen P. (2014) Cooperation, quorum sensing, and evolution of virulence in Staphylococcus aureus. Infection and Immunity, 82 (3). pp. 1045-1051. ISSN 1098-5522 http://iai.asm.org/content/82/3/1045 doi:10.1128/IAI.01216-13 doi:10.1128/IAI.01216-13
spellingShingle Pollitt, Eric J.G.
West, Stuart A.
Crusz, Shanika A.
Burton-Chellew, Maxwell N.
Diggle, Stephen P.
Cooperation, quorum sensing, and evolution of virulence in Staphylococcus aureus
title Cooperation, quorum sensing, and evolution of virulence in Staphylococcus aureus
title_full Cooperation, quorum sensing, and evolution of virulence in Staphylococcus aureus
title_fullStr Cooperation, quorum sensing, and evolution of virulence in Staphylococcus aureus
title_full_unstemmed Cooperation, quorum sensing, and evolution of virulence in Staphylococcus aureus
title_short Cooperation, quorum sensing, and evolution of virulence in Staphylococcus aureus
title_sort cooperation, quorum sensing, and evolution of virulence in staphylococcus aureus
url https://eprints.nottingham.ac.uk/2991/
https://eprints.nottingham.ac.uk/2991/
https://eprints.nottingham.ac.uk/2991/