A signature motif mediating selective interactions of BCL11A with the NR2E/F subfamily of orphan nuclear receptors

Despite their physiological importance, selective interactions between nuclear receptors (NRs) and their cofactors are poorly understood. Here, we describe a novel signature motif (F/YSXXLXXL/Y) in the developmental regulator BCL11A that facilitates its selective interaction with members of the NR2E...

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Main Authors: Chan, Chun Ming, Fulton, Joel, Montiel-Duarte, Cristina, Collins, Hilary M., Bharti, Neetu, Wadelin, Frances R., Moran, Paula M., Mongan, Nigel P., Heery, David M.
Format: Article
Published: Oxford University Press 2013
Online Access:https://eprints.nottingham.ac.uk/29027/
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author Chan, Chun Ming
Fulton, Joel
Montiel-Duarte, Cristina
Collins, Hilary M.
Bharti, Neetu
Wadelin, Frances R.
Moran, Paula M.
Mongan, Nigel P.
Heery, David M.
author_facet Chan, Chun Ming
Fulton, Joel
Montiel-Duarte, Cristina
Collins, Hilary M.
Bharti, Neetu
Wadelin, Frances R.
Moran, Paula M.
Mongan, Nigel P.
Heery, David M.
author_sort Chan, Chun Ming
building Nottingham Research Data Repository
collection Online Access
description Despite their physiological importance, selective interactions between nuclear receptors (NRs) and their cofactors are poorly understood. Here, we describe a novel signature motif (F/YSXXLXXL/Y) in the developmental regulator BCL11A that facilitates its selective interaction with members of the NR2E/F subfamily. Two copies of this motif (named here as RID1 and RID2) permit BCL11A to bind COUP-TFs (NR2F1;NR2F2;NR2F6) and Tailless/TLX (NR2E1), whereas RID1, but not RID2, binds PNR (NR2E3). We confirmed the existence of endogenous BCL11A/TLX complexes in mouse cortex tissue. No interactions of RID1 and RID2 with 20 other ligand-binding domains from different NR subtypes were observed. We show that RID1 and RID2 are required for BCL11A-mediated repression of endogenous γ-globin gene and the regulatory non-coding transcript Bgl3, and we identify COUP-TFII binding sites within the Bgl3 locus. In addition to their importance for BCL11A function, we show that F/YSXXLXXL/Y motifs are conserved in other NR cofactors. A single FSXXLXXL motif in the NR-binding SET domain protein NSD1 facilitates its interactions with the NR2E/F subfamily. However, the NSD1 motif incorporates features of both LXXLL and FSXXLXXL motifs, giving it a distinct NR-binding pattern in contrast to other cofactors. In summary, our results provide new insights into the selectivity of NR/cofactor complex formation.
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spelling nottingham-290272020-05-04T20:18:33Z https://eprints.nottingham.ac.uk/29027/ A signature motif mediating selective interactions of BCL11A with the NR2E/F subfamily of orphan nuclear receptors Chan, Chun Ming Fulton, Joel Montiel-Duarte, Cristina Collins, Hilary M. Bharti, Neetu Wadelin, Frances R. Moran, Paula M. Mongan, Nigel P. Heery, David M. Despite their physiological importance, selective interactions between nuclear receptors (NRs) and their cofactors are poorly understood. Here, we describe a novel signature motif (F/YSXXLXXL/Y) in the developmental regulator BCL11A that facilitates its selective interaction with members of the NR2E/F subfamily. Two copies of this motif (named here as RID1 and RID2) permit BCL11A to bind COUP-TFs (NR2F1;NR2F2;NR2F6) and Tailless/TLX (NR2E1), whereas RID1, but not RID2, binds PNR (NR2E3). We confirmed the existence of endogenous BCL11A/TLX complexes in mouse cortex tissue. No interactions of RID1 and RID2 with 20 other ligand-binding domains from different NR subtypes were observed. We show that RID1 and RID2 are required for BCL11A-mediated repression of endogenous γ-globin gene and the regulatory non-coding transcript Bgl3, and we identify COUP-TFII binding sites within the Bgl3 locus. In addition to their importance for BCL11A function, we show that F/YSXXLXXL/Y motifs are conserved in other NR cofactors. A single FSXXLXXL motif in the NR-binding SET domain protein NSD1 facilitates its interactions with the NR2E/F subfamily. However, the NSD1 motif incorporates features of both LXXLL and FSXXLXXL motifs, giving it a distinct NR-binding pattern in contrast to other cofactors. In summary, our results provide new insights into the selectivity of NR/cofactor complex formation. Oxford University Press 2013-11 Article PeerReviewed Chan, Chun Ming, Fulton, Joel, Montiel-Duarte, Cristina, Collins, Hilary M., Bharti, Neetu, Wadelin, Frances R., Moran, Paula M., Mongan, Nigel P. and Heery, David M. (2013) A signature motif mediating selective interactions of BCL11A with the NR2E/F subfamily of orphan nuclear receptors. Nucleic Acids Research, 41 (21). pp. 9663-9679. ISSN 1362-4962 http://nar.oxfordjournals.org/content/41/21/9663 doi:10.1093/nar/gkt761 doi:10.1093/nar/gkt761
spellingShingle Chan, Chun Ming
Fulton, Joel
Montiel-Duarte, Cristina
Collins, Hilary M.
Bharti, Neetu
Wadelin, Frances R.
Moran, Paula M.
Mongan, Nigel P.
Heery, David M.
A signature motif mediating selective interactions of BCL11A with the NR2E/F subfamily of orphan nuclear receptors
title A signature motif mediating selective interactions of BCL11A with the NR2E/F subfamily of orphan nuclear receptors
title_full A signature motif mediating selective interactions of BCL11A with the NR2E/F subfamily of orphan nuclear receptors
title_fullStr A signature motif mediating selective interactions of BCL11A with the NR2E/F subfamily of orphan nuclear receptors
title_full_unstemmed A signature motif mediating selective interactions of BCL11A with the NR2E/F subfamily of orphan nuclear receptors
title_short A signature motif mediating selective interactions of BCL11A with the NR2E/F subfamily of orphan nuclear receptors
title_sort signature motif mediating selective interactions of bcl11a with the nr2e/f subfamily of orphan nuclear receptors
url https://eprints.nottingham.ac.uk/29027/
https://eprints.nottingham.ac.uk/29027/
https://eprints.nottingham.ac.uk/29027/