DNA damage induces nucleoid compaction via the Mre11-Rad50 complex in the archaeon Haloferax volcanii

In prokaryotes the genome is organized in a dynamic structure called the nucleoid, which is embedded in the cytoplasm. We show here that in the archaeon Haloferax volcanii, compaction and reorganization of the nucleoid is induced by stresses that damage the genome or interfere with its replicat...

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Main Authors: Delmas, Stéphane, Duggin, Iain J., Allers, Thorsten
Format: Article
Published: Wiley 2013
Online Access:https://eprints.nottingham.ac.uk/2792/
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author Delmas, Stéphane
Duggin, Iain J.
Allers, Thorsten
author_facet Delmas, Stéphane
Duggin, Iain J.
Allers, Thorsten
author_sort Delmas, Stéphane
building Nottingham Research Data Repository
collection Online Access
description In prokaryotes the genome is organized in a dynamic structure called the nucleoid, which is embedded in the cytoplasm. We show here that in the archaeon Haloferax volcanii, compaction and reorganization of the nucleoid is induced by stresses that damage the genome or interfere with its replication. The fraction of cells exhibiting nucleoid compaction was proportional to the dose of the DNA damaging agent, and results obtained in cells defective for nucleotide excision repair suggest that breakage of DNA strands triggers reorganization of the nucleoid. We observed that compaction depends on the Mre11-Rad50 complex, suggesting a link to DNA double-strand break repair. However, compaction was observed in a radA mutant, indicating that the role of Mre11-Rad50 in nucleoid reorganisation is independent of homologous recombination. We therefore propose that nucleoid compaction is part of a DNA damage response that accelerates cell recovery by helping DNA repair proteins to locate their targets, and facilitating the search for intact DNA sequences during homologous recombination.
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spelling nottingham-27922020-05-04T20:19:40Z https://eprints.nottingham.ac.uk/2792/ DNA damage induces nucleoid compaction via the Mre11-Rad50 complex in the archaeon Haloferax volcanii Delmas, Stéphane Duggin, Iain J. Allers, Thorsten In prokaryotes the genome is organized in a dynamic structure called the nucleoid, which is embedded in the cytoplasm. We show here that in the archaeon Haloferax volcanii, compaction and reorganization of the nucleoid is induced by stresses that damage the genome or interfere with its replication. The fraction of cells exhibiting nucleoid compaction was proportional to the dose of the DNA damaging agent, and results obtained in cells defective for nucleotide excision repair suggest that breakage of DNA strands triggers reorganization of the nucleoid. We observed that compaction depends on the Mre11-Rad50 complex, suggesting a link to DNA double-strand break repair. However, compaction was observed in a radA mutant, indicating that the role of Mre11-Rad50 in nucleoid reorganisation is independent of homologous recombination. We therefore propose that nucleoid compaction is part of a DNA damage response that accelerates cell recovery by helping DNA repair proteins to locate their targets, and facilitating the search for intact DNA sequences during homologous recombination. Wiley 2013-01 Article PeerReviewed Delmas, Stéphane, Duggin, Iain J. and Allers, Thorsten (2013) DNA damage induces nucleoid compaction via the Mre11-Rad50 complex in the archaeon Haloferax volcanii. Molecular Microbiology, 87 (1). pp. 168-179. ISSN 0950-382X http://onlinelibrary.wiley.com/doi/10.1111/mmi.12091/abstract doi:10.1111/mmi.12091 doi:10.1111/mmi.12091
spellingShingle Delmas, Stéphane
Duggin, Iain J.
Allers, Thorsten
DNA damage induces nucleoid compaction via the Mre11-Rad50 complex in the archaeon Haloferax volcanii
title DNA damage induces nucleoid compaction via the Mre11-Rad50 complex in the archaeon Haloferax volcanii
title_full DNA damage induces nucleoid compaction via the Mre11-Rad50 complex in the archaeon Haloferax volcanii
title_fullStr DNA damage induces nucleoid compaction via the Mre11-Rad50 complex in the archaeon Haloferax volcanii
title_full_unstemmed DNA damage induces nucleoid compaction via the Mre11-Rad50 complex in the archaeon Haloferax volcanii
title_short DNA damage induces nucleoid compaction via the Mre11-Rad50 complex in the archaeon Haloferax volcanii
title_sort dna damage induces nucleoid compaction via the mre11-rad50 complex in the archaeon haloferax volcanii
url https://eprints.nottingham.ac.uk/2792/
https://eprints.nottingham.ac.uk/2792/
https://eprints.nottingham.ac.uk/2792/