Analysis of the Sam50 translocase of excavate organisms supports evolution of divergent organelles from a common endosymbiotic event

As free-living organisms the ancestors of mitochondria and plastids encoded complete genomes, proteomes and metabolomes. As these symbionts became organelles all these aspects were reduced – genomes have degenerated with the host nucleus now encoding the most of the remaining endosymbiont proteome,...

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Main Authors: Kay, Christopher J., Lawler, Karen, Kerr, Ian D.
Format: Article
Published: Portland Press: 2013
Online Access:https://eprints.nottingham.ac.uk/2686/
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author Kay, Christopher J.
Lawler, Karen
Kerr, Ian D.
author_facet Kay, Christopher J.
Lawler, Karen
Kerr, Ian D.
author_sort Kay, Christopher J.
building Nottingham Research Data Repository
collection Online Access
description As free-living organisms the ancestors of mitochondria and plastids encoded complete genomes, proteomes and metabolomes. As these symbionts became organelles all these aspects were reduced – genomes have degenerated with the host nucleus now encoding the most of the remaining endosymbiont proteome, while the metabolic processes of the symbiont have been streamlined to the functions of the emerging organelle. By contrast, the topology of the endosymbiont membrane has been preserved, necessitating the development of complex pathways for membrane insertion and translocation. In this study, we examine the characteristics of the endosymbiont-derived β-barrel insertase Sam501 in the excavate super-group. A candidate is further characterized in Trichomonas vaginalis, an unusual eukaryote possessing degenerate hydrogen-producing mitochondria called hydrogenosomes. This information supports a mitochondriate eukaryotic common ancestor with a similarly evolved β-barrel insertase, which has continued to be conserved in degenerate mitochondria.
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spelling nottingham-26862020-05-04T20:20:30Z https://eprints.nottingham.ac.uk/2686/ Analysis of the Sam50 translocase of excavate organisms supports evolution of divergent organelles from a common endosymbiotic event Kay, Christopher J. Lawler, Karen Kerr, Ian D. As free-living organisms the ancestors of mitochondria and plastids encoded complete genomes, proteomes and metabolomes. As these symbionts became organelles all these aspects were reduced – genomes have degenerated with the host nucleus now encoding the most of the remaining endosymbiont proteome, while the metabolic processes of the symbiont have been streamlined to the functions of the emerging organelle. By contrast, the topology of the endosymbiont membrane has been preserved, necessitating the development of complex pathways for membrane insertion and translocation. In this study, we examine the characteristics of the endosymbiont-derived β-barrel insertase Sam501 in the excavate super-group. A candidate is further characterized in Trichomonas vaginalis, an unusual eukaryote possessing degenerate hydrogen-producing mitochondria called hydrogenosomes. This information supports a mitochondriate eukaryotic common ancestor with a similarly evolved β-barrel insertase, which has continued to be conserved in degenerate mitochondria. Portland Press: 2013 Article PeerReviewed Kay, Christopher J., Lawler, Karen and Kerr, Ian D. (2013) Analysis of the Sam50 translocase of excavate organisms supports evolution of divergent organelles from a common endosymbiotic event. Bioscience Reports, 33 (6). e00084/1-e00084/9. ISSN 0144-8463 http://www.bioscirep.org/bsr/033/e084/bsr033e084.htm doi:10.1042/BSR20130049 doi:10.1042/BSR20130049
spellingShingle Kay, Christopher J.
Lawler, Karen
Kerr, Ian D.
Analysis of the Sam50 translocase of excavate organisms supports evolution of divergent organelles from a common endosymbiotic event
title Analysis of the Sam50 translocase of excavate organisms supports evolution of divergent organelles from a common endosymbiotic event
title_full Analysis of the Sam50 translocase of excavate organisms supports evolution of divergent organelles from a common endosymbiotic event
title_fullStr Analysis of the Sam50 translocase of excavate organisms supports evolution of divergent organelles from a common endosymbiotic event
title_full_unstemmed Analysis of the Sam50 translocase of excavate organisms supports evolution of divergent organelles from a common endosymbiotic event
title_short Analysis of the Sam50 translocase of excavate organisms supports evolution of divergent organelles from a common endosymbiotic event
title_sort analysis of the sam50 translocase of excavate organisms supports evolution of divergent organelles from a common endosymbiotic event
url https://eprints.nottingham.ac.uk/2686/
https://eprints.nottingham.ac.uk/2686/
https://eprints.nottingham.ac.uk/2686/