Travelling waves in a neural field model with refractoriness

At one level of abstraction neural tissue can be regarded as a medium for turning local synaptic activity into output signals that propagate over large distances via axons to generate further synaptic activity that can cause reverberant activity in networks that possess a mixture of excitatory and i...

Full description

Bibliographic Details
Main Authors: Meijer, Hil G.E., Coombes, Stephen
Format: Article
Published: Springer Verlag 2014
Subjects:
Online Access:https://eprints.nottingham.ac.uk/2670/
_version_ 1848790845305651200
author Meijer, Hil G.E.
Coombes, Stephen
author_facet Meijer, Hil G.E.
Coombes, Stephen
author_sort Meijer, Hil G.E.
building Nottingham Research Data Repository
collection Online Access
description At one level of abstraction neural tissue can be regarded as a medium for turning local synaptic activity into output signals that propagate over large distances via axons to generate further synaptic activity that can cause reverberant activity in networks that possess a mixture of excitatory and inhibitory connections. This output is often taken to be a firing rate, and the mathematical form for the evolution equation of activity depends upon a spatial convolution of this rate with a fixed anatomical connectivity pattern. Such formulations often neglect the metabolic processes that would ultimately limit synaptic activity. Here we reinstate such a process, in the spirit of an original prescription by Wilson and Cowan (Biophys J 12:1–24, 1972 ), using a term that multiplies the usual spatial convolution with a moving time average of local activity over some refractory time-scale. This modulation can substantially affect network behaviour, and in particular give rise to periodic travelling waves in a purely excitatory network (with exponentially decaying anatomical connectivity), which in the absence of refractoriness would only support travelling fronts.We construct these solutions numerically as stationary periodic solutions in a co-moving frame (of both an equivalent delay differential model as well as the original delay integro-differential model). Continuation methods are used to obtain the dispersion curve for periodic travelling waves (speed as a function of period), and found to be reminiscent of those for spatially extended models of excitable tissue. A kinematic analysis (based on the dispersion curve) predicts the onset of wave instabilities, which are confirmed numerically.
first_indexed 2025-11-14T18:19:05Z
format Article
id nottingham-2670
institution University of Nottingham Malaysia Campus
institution_category Local University
last_indexed 2025-11-14T18:19:05Z
publishDate 2014
publisher Springer Verlag
recordtype eprints
repository_type Digital Repository
spelling nottingham-26702020-05-04T16:43:46Z https://eprints.nottingham.ac.uk/2670/ Travelling waves in a neural field model with refractoriness Meijer, Hil G.E. Coombes, Stephen At one level of abstraction neural tissue can be regarded as a medium for turning local synaptic activity into output signals that propagate over large distances via axons to generate further synaptic activity that can cause reverberant activity in networks that possess a mixture of excitatory and inhibitory connections. This output is often taken to be a firing rate, and the mathematical form for the evolution equation of activity depends upon a spatial convolution of this rate with a fixed anatomical connectivity pattern. Such formulations often neglect the metabolic processes that would ultimately limit synaptic activity. Here we reinstate such a process, in the spirit of an original prescription by Wilson and Cowan (Biophys J 12:1–24, 1972 ), using a term that multiplies the usual spatial convolution with a moving time average of local activity over some refractory time-scale. This modulation can substantially affect network behaviour, and in particular give rise to periodic travelling waves in a purely excitatory network (with exponentially decaying anatomical connectivity), which in the absence of refractoriness would only support travelling fronts.We construct these solutions numerically as stationary periodic solutions in a co-moving frame (of both an equivalent delay differential model as well as the original delay integro-differential model). Continuation methods are used to obtain the dispersion curve for periodic travelling waves (speed as a function of period), and found to be reminiscent of those for spatially extended models of excitable tissue. A kinematic analysis (based on the dispersion curve) predicts the onset of wave instabilities, which are confirmed numerically. Springer Verlag 2014-04-01 Article PeerReviewed Meijer, Hil G.E. and Coombes, Stephen (2014) Travelling waves in a neural field model with refractoriness. Journal of Mathematical Biology, 68 (5). pp. 1249-1268. ISSN 1432-1416 neural field models; travelling waves; refractoriness; delay differential equations http://link.springer.com/article/10.1007%2Fs00285-013-0670-x doi:10.1007/s00285-013-0670-x doi:10.1007/s00285-013-0670-x
spellingShingle neural field models; travelling waves; refractoriness; delay differential equations
Meijer, Hil G.E.
Coombes, Stephen
Travelling waves in a neural field model with refractoriness
title Travelling waves in a neural field model with refractoriness
title_full Travelling waves in a neural field model with refractoriness
title_fullStr Travelling waves in a neural field model with refractoriness
title_full_unstemmed Travelling waves in a neural field model with refractoriness
title_short Travelling waves in a neural field model with refractoriness
title_sort travelling waves in a neural field model with refractoriness
topic neural field models; travelling waves; refractoriness; delay differential equations
url https://eprints.nottingham.ac.uk/2670/
https://eprints.nottingham.ac.uk/2670/
https://eprints.nottingham.ac.uk/2670/