The transposon-like correia elements encode numerous strong promoters and provide a potential new mechanism for phase variation in the meningococcus

Neisseria meningitidis is the primary causative agent of bacterial meningitis. The genome is rich in repetitive DNA and almost 2% is occupied by a diminutive transposon called the Correia element. Here we report a bioinformatic analysis defining eight subtypes of the element with four distinct types...

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Main Authors: Siddique, Azeem, Buisine, Nicolas, Chalmers, Ronald
Format: Article
Published: Public Library of Science 2011
Online Access:https://eprints.nottingham.ac.uk/2471/
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author Siddique, Azeem
Buisine, Nicolas
Chalmers, Ronald
author_facet Siddique, Azeem
Buisine, Nicolas
Chalmers, Ronald
author_sort Siddique, Azeem
building Nottingham Research Data Repository
collection Online Access
description Neisseria meningitidis is the primary causative agent of bacterial meningitis. The genome is rich in repetitive DNA and almost 2% is occupied by a diminutive transposon called the Correia element. Here we report a bioinformatic analysis defining eight subtypes of the element with four distinct types of ends. Transcriptional analysis, using PCR and a lacZ reporter system, revealed that two ends in particular encode strong promoters. The activity of the strongest promoter is dictated by a recurrent polymorphism (Y128) at the right end of the element. We highlight examples of elements that appear to drive transcription of adjacent genes and others that may express small non-coding RNAs. Pair-wise comparisons between three meningococcal genomes revealed that no more than two-thirds of Correia elements maintain their subtype at any particular locus. This is due to recombinational class switching between elements in a single strain. Upon switching subtype, a new allele is available to spread through the population by natural transformation. This process may represent a hitherto unrecognized mechanism for phase variation in the meningococcus. We conclude that the strain-to-strain variability of the Correia elements, and the large number of strong promoters encoded by them, allows for potentially widespread effects within the population as a whole. By defining the strength of the promoters encoded by the eight subtypes of Correia ends, we provide a resource that allows the transcriptional effects of a particular subtype at a given locus to be predicted.
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spelling nottingham-24712020-05-04T16:30:15Z https://eprints.nottingham.ac.uk/2471/ The transposon-like correia elements encode numerous strong promoters and provide a potential new mechanism for phase variation in the meningococcus Siddique, Azeem Buisine, Nicolas Chalmers, Ronald Neisseria meningitidis is the primary causative agent of bacterial meningitis. The genome is rich in repetitive DNA and almost 2% is occupied by a diminutive transposon called the Correia element. Here we report a bioinformatic analysis defining eight subtypes of the element with four distinct types of ends. Transcriptional analysis, using PCR and a lacZ reporter system, revealed that two ends in particular encode strong promoters. The activity of the strongest promoter is dictated by a recurrent polymorphism (Y128) at the right end of the element. We highlight examples of elements that appear to drive transcription of adjacent genes and others that may express small non-coding RNAs. Pair-wise comparisons between three meningococcal genomes revealed that no more than two-thirds of Correia elements maintain their subtype at any particular locus. This is due to recombinational class switching between elements in a single strain. Upon switching subtype, a new allele is available to spread through the population by natural transformation. This process may represent a hitherto unrecognized mechanism for phase variation in the meningococcus. We conclude that the strain-to-strain variability of the Correia elements, and the large number of strong promoters encoded by them, allows for potentially widespread effects within the population as a whole. By defining the strength of the promoters encoded by the eight subtypes of Correia ends, we provide a resource that allows the transcriptional effects of a particular subtype at a given locus to be predicted. Public Library of Science 2011-01-20 Article PeerReviewed Siddique, Azeem, Buisine, Nicolas and Chalmers, Ronald (2011) The transposon-like correia elements encode numerous strong promoters and provide a potential new mechanism for phase variation in the meningococcus. PLoS Genetics, 7 (1). 13/1-13/13. ISSN 1553-7390 http://www.plosgenetics.org/article/info%3Adoi%2F10.1371%2Fjournal.pgen.1001277 doi:10.1371/journal.pgen.1001277 doi:10.1371/journal.pgen.1001277
spellingShingle Siddique, Azeem
Buisine, Nicolas
Chalmers, Ronald
The transposon-like correia elements encode numerous strong promoters and provide a potential new mechanism for phase variation in the meningococcus
title The transposon-like correia elements encode numerous strong promoters and provide a potential new mechanism for phase variation in the meningococcus
title_full The transposon-like correia elements encode numerous strong promoters and provide a potential new mechanism for phase variation in the meningococcus
title_fullStr The transposon-like correia elements encode numerous strong promoters and provide a potential new mechanism for phase variation in the meningococcus
title_full_unstemmed The transposon-like correia elements encode numerous strong promoters and provide a potential new mechanism for phase variation in the meningococcus
title_short The transposon-like correia elements encode numerous strong promoters and provide a potential new mechanism for phase variation in the meningococcus
title_sort transposon-like correia elements encode numerous strong promoters and provide a potential new mechanism for phase variation in the meningococcus
url https://eprints.nottingham.ac.uk/2471/
https://eprints.nottingham.ac.uk/2471/
https://eprints.nottingham.ac.uk/2471/