Investigating a conserved role for Nanog in primordial germ cell specification

The specification of Primordial Germ Cells (PGCs) in early mammalian development is a fundamental process of development, critical for, maintaining the germ line. There are two methods by which PGCs are specified in the animal kingdom. Firstly, through the predetermined method involving maternally i...

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Main Author: Shephard, Matthew Thomas
Format: Thesis (University of Nottingham only)
Language:English
Published: 2010
Online Access:https://eprints.nottingham.ac.uk/11663/
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author Shephard, Matthew Thomas
author_facet Shephard, Matthew Thomas
author_sort Shephard, Matthew Thomas
building Nottingham Research Data Repository
collection Online Access
description The specification of Primordial Germ Cells (PGCs) in early mammalian development is a fundamental process of development, critical for, maintaining the germ line. There are two methods by which PGCs are specified in the animal kingdom. Firstly, through the predetermined method involving maternally inherited germplasm, and secondly through the induction of pluripotent precursors in a process termed epigenesis. Embryoid Bodies (EBs) can be used to recapitulate early developmental processes of mammalian development, allowing the dissection and better understanding of critical processes. This project will aim to try and better understand the role of the homeobox transcription factor Nanog in specifying PGCs through the process of epigenesis. Mouse Nanog over-expression in EBs has been shown to increase PGC specification, assayed through RT-PCR analysis. The functional activity of ancestral Nanog is hypothesised to be conserved through the major trunk of evolution in organisms that specify PGCs through the induced method of epigenesis. To test this hypothesis, ancient Nanog orthologs were isolated from different positions of the Deuterostome superfamily and over-expressed in EBs. Their effects on enhancing PGC specification were then quantified through quantitative RT-PCR and western blot analysis. The results from this project show that the over-expression of ancient Nanog orthologs in mES cells can enhance the process of PGC specification, by increasing the PGC associated markers Dazl and Vasa. This suggests a possible conservation of functional Nanog activity in organisms that specify PGCs through epigenesis, and also that pluripotency is an ancient trait.
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spelling nottingham-116632025-02-28T11:14:52Z https://eprints.nottingham.ac.uk/11663/ Investigating a conserved role for Nanog in primordial germ cell specification Shephard, Matthew Thomas The specification of Primordial Germ Cells (PGCs) in early mammalian development is a fundamental process of development, critical for, maintaining the germ line. There are two methods by which PGCs are specified in the animal kingdom. Firstly, through the predetermined method involving maternally inherited germplasm, and secondly through the induction of pluripotent precursors in a process termed epigenesis. Embryoid Bodies (EBs) can be used to recapitulate early developmental processes of mammalian development, allowing the dissection and better understanding of critical processes. This project will aim to try and better understand the role of the homeobox transcription factor Nanog in specifying PGCs through the process of epigenesis. Mouse Nanog over-expression in EBs has been shown to increase PGC specification, assayed through RT-PCR analysis. The functional activity of ancestral Nanog is hypothesised to be conserved through the major trunk of evolution in organisms that specify PGCs through the induced method of epigenesis. To test this hypothesis, ancient Nanog orthologs were isolated from different positions of the Deuterostome superfamily and over-expressed in EBs. Their effects on enhancing PGC specification were then quantified through quantitative RT-PCR and western blot analysis. The results from this project show that the over-expression of ancient Nanog orthologs in mES cells can enhance the process of PGC specification, by increasing the PGC associated markers Dazl and Vasa. This suggests a possible conservation of functional Nanog activity in organisms that specify PGCs through epigenesis, and also that pluripotency is an ancient trait. 2010-12-10 Thesis (University of Nottingham only) NonPeerReviewed application/pdf en arr https://eprints.nottingham.ac.uk/11663/2/Matthew_Shephard_E-Thesis.pdf Shephard, Matthew Thomas (2010) Investigating a conserved role for Nanog in primordial germ cell specification. MRes thesis, University of Nottingham.
spellingShingle Shephard, Matthew Thomas
Investigating a conserved role for Nanog in primordial germ cell specification
title Investigating a conserved role for Nanog in primordial germ cell specification
title_full Investigating a conserved role for Nanog in primordial germ cell specification
title_fullStr Investigating a conserved role for Nanog in primordial germ cell specification
title_full_unstemmed Investigating a conserved role for Nanog in primordial germ cell specification
title_short Investigating a conserved role for Nanog in primordial germ cell specification
title_sort investigating a conserved role for nanog in primordial germ cell specification
url https://eprints.nottingham.ac.uk/11663/