Seed-coat thickness explains contrasting germination responses to smoke and heat in Leucadendron

Fire stimulates the germination of most seeds in fire-prone vegetation. Fruits of Leucadendron (Proteaceae) are winged achenes or nutlets that correlate with their requirements for smoke and/or heat in promoting germination. We describe five possible smoke–heat dormancy-release/germination syndromes...

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Main Authors: Lamont, Byron, Gomez Barreiro, Pablo, Newton, Rosemary J
Format: Journal Article
Language:English
Published: CAMBRIDGE UNIV PRESS 2022
Subjects:
Online Access:http://purl.org/au-research/grants/arc/DP130103029
http://hdl.handle.net/20.500.11937/90941
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author Lamont, Byron
Gomez Barreiro, Pablo
Newton, Rosemary J
author_facet Lamont, Byron
Gomez Barreiro, Pablo
Newton, Rosemary J
author_sort Lamont, Byron
building Curtin Institutional Repository
collection Online Access
description Fire stimulates the germination of most seeds in fire-prone vegetation. Fruits of Leucadendron (Proteaceae) are winged achenes or nutlets that correlate with their requirements for smoke and/or heat in promoting germination. We describe five possible smoke–heat dormancy-release/germination syndromes among plants, of which Leucadendron displays three (no response, smoke only, smoke and heat). As seed-coat thickness varies with seed-storage location (plant or soil) and morphology (winged or wingless), we tested its possible role in water uptake and germination. Species with winged seeds achieved 100% germination in the absence of smoke/heat, seed coats were an order of magnitude thinner, and their permeability greatly exceeded that of nutlets. As seed-coat thickness increased (1) imbibitional water uptake declined at a decreasing rate, and (2) the response to smoke, and to a lesser extent heat, increased linearly to reach levels of germination approaching those of winged seeds. For species responsive to smoke and heat, there was no additive effect when applied together, suggesting that they may have promoted the same physiological process. We conclude that seed-coat thickness holds the key to germination requirements in this genus, independent of seed-storage location or morphology. By what mechanisms (1) the smoke response is greater the thicker the seed coat and (2) smoke chemicals might increase water permeability to explain the non-additive effect of smoke and heat, warrant further investigation.
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spelling curtin-20.500.11937-909412023-05-11T07:17:32Z Seed-coat thickness explains contrasting germination responses to smoke and heat in Leucadendron Lamont, Byron Gomez Barreiro, Pablo Newton, Rosemary J Science & Technology Life Sciences & Biomedicine Plant Sciences heat Leucadendron Mediterranean-type ecosystems seed-coat thickness and permeability serotiny smoke soil-stored seeds FIRE WATER PROTEACEAE CUES KARRIKINOLIDE TEMPERATURE DISPERSAL STIMULATE FRUITS Fire stimulates the germination of most seeds in fire-prone vegetation. Fruits of Leucadendron (Proteaceae) are winged achenes or nutlets that correlate with their requirements for smoke and/or heat in promoting germination. We describe five possible smoke–heat dormancy-release/germination syndromes among plants, of which Leucadendron displays three (no response, smoke only, smoke and heat). As seed-coat thickness varies with seed-storage location (plant or soil) and morphology (winged or wingless), we tested its possible role in water uptake and germination. Species with winged seeds achieved 100% germination in the absence of smoke/heat, seed coats were an order of magnitude thinner, and their permeability greatly exceeded that of nutlets. As seed-coat thickness increased (1) imbibitional water uptake declined at a decreasing rate, and (2) the response to smoke, and to a lesser extent heat, increased linearly to reach levels of germination approaching those of winged seeds. For species responsive to smoke and heat, there was no additive effect when applied together, suggesting that they may have promoted the same physiological process. We conclude that seed-coat thickness holds the key to germination requirements in this genus, independent of seed-storage location or morphology. By what mechanisms (1) the smoke response is greater the thicker the seed coat and (2) smoke chemicals might increase water permeability to explain the non-additive effect of smoke and heat, warrant further investigation. 2022 Journal Article http://hdl.handle.net/20.500.11937/90941 10.1017/S0960258522000113 English http://purl.org/au-research/grants/arc/DP130103029 http://creativecommons.org/licenses/by/4.0/ CAMBRIDGE UNIV PRESS fulltext
spellingShingle Science & Technology
Life Sciences & Biomedicine
Plant Sciences
heat
Leucadendron
Mediterranean-type ecosystems
seed-coat thickness and permeability
serotiny
smoke
soil-stored seeds
FIRE
WATER
PROTEACEAE
CUES
KARRIKINOLIDE
TEMPERATURE
DISPERSAL
STIMULATE
FRUITS
Lamont, Byron
Gomez Barreiro, Pablo
Newton, Rosemary J
Seed-coat thickness explains contrasting germination responses to smoke and heat in Leucadendron
title Seed-coat thickness explains contrasting germination responses to smoke and heat in Leucadendron
title_full Seed-coat thickness explains contrasting germination responses to smoke and heat in Leucadendron
title_fullStr Seed-coat thickness explains contrasting germination responses to smoke and heat in Leucadendron
title_full_unstemmed Seed-coat thickness explains contrasting germination responses to smoke and heat in Leucadendron
title_short Seed-coat thickness explains contrasting germination responses to smoke and heat in Leucadendron
title_sort seed-coat thickness explains contrasting germination responses to smoke and heat in leucadendron
topic Science & Technology
Life Sciences & Biomedicine
Plant Sciences
heat
Leucadendron
Mediterranean-type ecosystems
seed-coat thickness and permeability
serotiny
smoke
soil-stored seeds
FIRE
WATER
PROTEACEAE
CUES
KARRIKINOLIDE
TEMPERATURE
DISPERSAL
STIMULATE
FRUITS
url http://purl.org/au-research/grants/arc/DP130103029
http://hdl.handle.net/20.500.11937/90941