Transcription factor lineages in plant-pathogenic fungi, connecting diversity with fungal virulence

Plant-pathogenic fungi span diverse taxonomic lineages. Their host-infection strategies are often specialised and require the coordinated regulation of molecular virulence factors. Transcription factors (TFs) are fundamental regulators of gene expression, yet relatively few virulence-specific regula...

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Main Authors: John, Evan, Singh, Karam, Oliver, Richard, Tan, Kar-Chun
Format: Journal Article
Published: Elsevier 2022
Online Access:http://hdl.handle.net/20.500.11937/88799
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author John, Evan
Singh, Karam
Oliver, Richard
Tan, Kar-Chun
author_facet John, Evan
Singh, Karam
Oliver, Richard
Tan, Kar-Chun
author_sort John, Evan
building Curtin Institutional Repository
collection Online Access
description Plant-pathogenic fungi span diverse taxonomic lineages. Their host-infection strategies are often specialised and require the coordinated regulation of molecular virulence factors. Transcription factors (TFs) are fundamental regulators of gene expression, yet relatively few virulence-specific regulators are characterised in detail and their evolutionary trajectories are not well understood. Hence, this study compared the full range of TFs across taxonomically-diverse fungal proteomes and classified their lineages through an orthology analysis. The primary aims were to characterise differences in the range and profile of TF lineages broadly linked to plant-host association or pathogenic lifestyles, and to better characterise the evolutionary origin and trajectory of experimentally-validated virulence regulators. We observed significantly fewer TFs among obligate, host-associated pathogens, largely attributed to contractions in several Zn2Cys6 TF-orthogroup lineages. We also present novel insight into the key virulence-regulating TFs Ste12, Pf2 and EBR1, providing evidence for their ancestral origins, expansion and/or loss. Ultimately, the analysis presented here provides both primary evidence for TF evolution in fungal phytopathogenicity, as well as a practical phylogenetic resource to guide further detailed investigation on the regulation of virulence within key pathogen lineages.
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spelling curtin-20.500.11937-887992022-07-07T08:31:15Z Transcription factor lineages in plant-pathogenic fungi, connecting diversity with fungal virulence John, Evan Singh, Karam Oliver, Richard Tan, Kar-Chun Plant-pathogenic fungi span diverse taxonomic lineages. Their host-infection strategies are often specialised and require the coordinated regulation of molecular virulence factors. Transcription factors (TFs) are fundamental regulators of gene expression, yet relatively few virulence-specific regulators are characterised in detail and their evolutionary trajectories are not well understood. Hence, this study compared the full range of TFs across taxonomically-diverse fungal proteomes and classified their lineages through an orthology analysis. The primary aims were to characterise differences in the range and profile of TF lineages broadly linked to plant-host association or pathogenic lifestyles, and to better characterise the evolutionary origin and trajectory of experimentally-validated virulence regulators. We observed significantly fewer TFs among obligate, host-associated pathogens, largely attributed to contractions in several Zn2Cys6 TF-orthogroup lineages. We also present novel insight into the key virulence-regulating TFs Ste12, Pf2 and EBR1, providing evidence for their ancestral origins, expansion and/or loss. Ultimately, the analysis presented here provides both primary evidence for TF evolution in fungal phytopathogenicity, as well as a practical phylogenetic resource to guide further detailed investigation on the regulation of virulence within key pathogen lineages. 2022 Journal Article http://hdl.handle.net/20.500.11937/88799 10.1016/j.fgb.2022.103712 http://creativecommons.org/licenses/by/4.0/ Elsevier fulltext
spellingShingle John, Evan
Singh, Karam
Oliver, Richard
Tan, Kar-Chun
Transcription factor lineages in plant-pathogenic fungi, connecting diversity with fungal virulence
title Transcription factor lineages in plant-pathogenic fungi, connecting diversity with fungal virulence
title_full Transcription factor lineages in plant-pathogenic fungi, connecting diversity with fungal virulence
title_fullStr Transcription factor lineages in plant-pathogenic fungi, connecting diversity with fungal virulence
title_full_unstemmed Transcription factor lineages in plant-pathogenic fungi, connecting diversity with fungal virulence
title_short Transcription factor lineages in plant-pathogenic fungi, connecting diversity with fungal virulence
title_sort transcription factor lineages in plant-pathogenic fungi, connecting diversity with fungal virulence
url http://hdl.handle.net/20.500.11937/88799