Investigating the role of calcium/calmodulin-dependent protein kinases in Stagonospora nodorum

Three genes encoding different Ca2+/calmodulin-dependent protein kinases have been characterized in the wheat phytopathogenic fungus Stagonospora nodorum. The kinases were identified from the S. nodorum genome sequence on the basis of sequence homology to known Ca2+/calmodulin-dependent protein kina...

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Main Authors: Solomon, P., Rybak, K., Trengove, R., Oliver, Richard
Format: Journal Article
Published: BLACKWELL PUBLISHING 2006
Subjects:
Online Access:http://hdl.handle.net/20.500.11937/42967
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author Solomon, P.
Rybak, K.
Trengove, R.
Oliver, Richard
author_facet Solomon, P.
Rybak, K.
Trengove, R.
Oliver, Richard
author_sort Solomon, P.
building Curtin Institutional Repository
collection Online Access
description Three genes encoding different Ca2+/calmodulin-dependent protein kinases have been characterized in the wheat phytopathogenic fungus Stagonospora nodorum. The kinases were identified from the S. nodorum genome sequence on the basis of sequence homology to known Ca2+/calmodulin-dependent protein kinases. Expression analysis determined that each of the kinases was expressed during growth in vitro and also during infection. The onset of sporulation triggered increased transcript levels of each of the kinases, particularly CpkA where an 11-fold increase in expression was observed during sporulation in planta. The role of the kinases was further determined via a reverse genetics approach. The disruption of CpkA affected vegetative growth in vitro and also sporulation. The cpkA strains produced 20-fold less spores on complex media and were unable to sporulate on defined minimal media. Infection assays showed that CpkA was not required for lesion development but was essential for sporulation at the completion of the infection cycle. Microscopic analysis revealed that the disruption of CpkA resulted in Stagonospora nodorum being unable to differentiate the mycelial knot into immature pycnidia during sporulation. A metabolite analysis of infected leaves during sporulation excluded the possible involvement of mannitol, a compound previously shown to be involved in the sporulation of Stagonospora nodorum. The disruption of CpkB did not effect growth in vitro or pathogenicity. Stagonospora nodorum strains lacking CpkC appeared unaffected during growth in planta but showed delayed lesion development and sporulation during infection.
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spelling curtin-20.500.11937-429672019-05-21T07:31:53Z Investigating the role of calcium/calmodulin-dependent protein kinases in Stagonospora nodorum Solomon, P. Rybak, K. Trengove, R. Oliver, Richard GENE STAINING TECHNIQUE NUCLEAR DIVISION GROWTH CRYPTOCOCCUS-NEOFORMANS EXPRESSION PATHOGEN CLADOSPORIUM-FULVUM CA2+/CALMODULIN-DEPENDENT KINASE ASPERGILLUS-NIDULANS WHEAT Three genes encoding different Ca2+/calmodulin-dependent protein kinases have been characterized in the wheat phytopathogenic fungus Stagonospora nodorum. The kinases were identified from the S. nodorum genome sequence on the basis of sequence homology to known Ca2+/calmodulin-dependent protein kinases. Expression analysis determined that each of the kinases was expressed during growth in vitro and also during infection. The onset of sporulation triggered increased transcript levels of each of the kinases, particularly CpkA where an 11-fold increase in expression was observed during sporulation in planta. The role of the kinases was further determined via a reverse genetics approach. The disruption of CpkA affected vegetative growth in vitro and also sporulation. The cpkA strains produced 20-fold less spores on complex media and were unable to sporulate on defined minimal media. Infection assays showed that CpkA was not required for lesion development but was essential for sporulation at the completion of the infection cycle. Microscopic analysis revealed that the disruption of CpkA resulted in Stagonospora nodorum being unable to differentiate the mycelial knot into immature pycnidia during sporulation. A metabolite analysis of infected leaves during sporulation excluded the possible involvement of mannitol, a compound previously shown to be involved in the sporulation of Stagonospora nodorum. The disruption of CpkB did not effect growth in vitro or pathogenicity. Stagonospora nodorum strains lacking CpkC appeared unaffected during growth in planta but showed delayed lesion development and sporulation during infection. 2006 Journal Article http://hdl.handle.net/20.500.11937/42967 10.1111/j.1365-2958.2006.05380.x BLACKWELL PUBLISHING restricted
spellingShingle GENE
STAINING TECHNIQUE
NUCLEAR DIVISION
GROWTH
CRYPTOCOCCUS-NEOFORMANS
EXPRESSION
PATHOGEN CLADOSPORIUM-FULVUM
CA2+/CALMODULIN-DEPENDENT KINASE
ASPERGILLUS-NIDULANS
WHEAT
Solomon, P.
Rybak, K.
Trengove, R.
Oliver, Richard
Investigating the role of calcium/calmodulin-dependent protein kinases in Stagonospora nodorum
title Investigating the role of calcium/calmodulin-dependent protein kinases in Stagonospora nodorum
title_full Investigating the role of calcium/calmodulin-dependent protein kinases in Stagonospora nodorum
title_fullStr Investigating the role of calcium/calmodulin-dependent protein kinases in Stagonospora nodorum
title_full_unstemmed Investigating the role of calcium/calmodulin-dependent protein kinases in Stagonospora nodorum
title_short Investigating the role of calcium/calmodulin-dependent protein kinases in Stagonospora nodorum
title_sort investigating the role of calcium/calmodulin-dependent protein kinases in stagonospora nodorum
topic GENE
STAINING TECHNIQUE
NUCLEAR DIVISION
GROWTH
CRYPTOCOCCUS-NEOFORMANS
EXPRESSION
PATHOGEN CLADOSPORIUM-FULVUM
CA2+/CALMODULIN-DEPENDENT KINASE
ASPERGILLUS-NIDULANS
WHEAT
url http://hdl.handle.net/20.500.11937/42967