A 350-million-year legacy of fire adaptation among conifers

Current phylogenetic evidence shows that fire began shaping the evolution of land plants 125 Ma, although the fossil charcoal record indicates that fire has a much longer history (>350 Ma). Serotiny (on-plant seed storage) is generally accepted as an adaptation to fire among woody plants. We...

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Main Authors: He, Tianhua, Belcher, C., Lamont, Byron, Lim, S.
Format: Journal Article
Published: 2015
Online Access:http://purl.org/au-research/grants/arc/DP120103389
http://hdl.handle.net/20.500.11937/16646
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author He, Tianhua
Belcher, C.
Lamont, Byron
Lim, S.
author_facet He, Tianhua
Belcher, C.
Lamont, Byron
Lim, S.
author_sort He, Tianhua
building Curtin Institutional Repository
collection Online Access
description Current phylogenetic evidence shows that fire began shaping the evolution of land plants 125 Ma, although the fossil charcoal record indicates that fire has a much longer history (>350 Ma). Serotiny (on-plant seed storage) is generally accepted as an adaptation to fire among woody plants. We developed a conceptual model of the requirements for the evolution of serotiny, and propose that serotiny is only expressed in the presence of a woody rachis as supporting structure, compact scales covering seeds as protective structure, seed wing as dispersal structure, and crown fire as the agent of selection and mechanism for seed release. This model is strongly supported by empirical data for modern ecosystems. We reconstructed the evolutionary history of intrinsic structural states required for the expression of serotiny in conifers, and show that these were diagnostic for early ('transitional') conifers from 332 Ma (late-Carboniferous). We assessed the likely flammable characteristics of early conifers and found that scale-leaved conifers burn rapidly and with high intensity, supporting the idea that crown fire regimes may have dominated early conifer ecosystems. Synthesis. Coupled with strong evidence for frequent fire throughout the Permian-Carboniferous and fossil evidence for other fire-related traits, we conclude that many early conifers were serotinous in response to intense crown fires, indicating that fire may have had a major impact on the evolution of plant traits as far back as 350 Ma.
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spelling curtin-20.500.11937-166462019-02-19T05:34:59Z A 350-million-year legacy of fire adaptation among conifers He, Tianhua Belcher, C. Lamont, Byron Lim, S. Current phylogenetic evidence shows that fire began shaping the evolution of land plants 125 Ma, although the fossil charcoal record indicates that fire has a much longer history (>350 Ma). Serotiny (on-plant seed storage) is generally accepted as an adaptation to fire among woody plants. We developed a conceptual model of the requirements for the evolution of serotiny, and propose that serotiny is only expressed in the presence of a woody rachis as supporting structure, compact scales covering seeds as protective structure, seed wing as dispersal structure, and crown fire as the agent of selection and mechanism for seed release. This model is strongly supported by empirical data for modern ecosystems. We reconstructed the evolutionary history of intrinsic structural states required for the expression of serotiny in conifers, and show that these were diagnostic for early ('transitional') conifers from 332 Ma (late-Carboniferous). We assessed the likely flammable characteristics of early conifers and found that scale-leaved conifers burn rapidly and with high intensity, supporting the idea that crown fire regimes may have dominated early conifer ecosystems. Synthesis. Coupled with strong evidence for frequent fire throughout the Permian-Carboniferous and fossil evidence for other fire-related traits, we conclude that many early conifers were serotinous in response to intense crown fires, indicating that fire may have had a major impact on the evolution of plant traits as far back as 350 Ma. 2015 Journal Article http://hdl.handle.net/20.500.11937/16646 10.1111/1365-2745.12513 http://purl.org/au-research/grants/arc/DP120103389 fulltext
spellingShingle He, Tianhua
Belcher, C.
Lamont, Byron
Lim, S.
A 350-million-year legacy of fire adaptation among conifers
title A 350-million-year legacy of fire adaptation among conifers
title_full A 350-million-year legacy of fire adaptation among conifers
title_fullStr A 350-million-year legacy of fire adaptation among conifers
title_full_unstemmed A 350-million-year legacy of fire adaptation among conifers
title_short A 350-million-year legacy of fire adaptation among conifers
title_sort 350-million-year legacy of fire adaptation among conifers
url http://purl.org/au-research/grants/arc/DP120103389
http://hdl.handle.net/20.500.11937/16646