Unique Biochemical and Mineral Composition of Whale Ear Bones

Cetaceans are obligate aquatic mammals derived from terrestrial artiodactyls. The defining characteristic of cetaceans is a thick and dense lip (pachyosteosclerotic involucrum) of an ear bone (the tympanic). This unique feature is absent in modern terrestrial artiodactyls and is suggested to be impo...

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Main Authors: Kim, S., Thewissen, J., Churchill, M., Suydam, R., Ketten, Darlene, Clementz, M.
Format: Journal Article
Published: The University of Chicago Press 2014
Subjects:
Online Access:http://hdl.handle.net/20.500.11937/11090
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author Kim, S.
Thewissen, J.
Churchill, M.
Suydam, R.
Ketten, Darlene
Clementz, M.
author_facet Kim, S.
Thewissen, J.
Churchill, M.
Suydam, R.
Ketten, Darlene
Clementz, M.
author_sort Kim, S.
building Curtin Institutional Repository
collection Online Access
description Cetaceans are obligate aquatic mammals derived from terrestrial artiodactyls. The defining characteristic of cetaceans is a thick and dense lip (pachyosteosclerotic involucrum) of an ear bone (the tympanic). This unique feature is absent in modern terrestrial artiodactyls and is suggested to be important in underwater hearing. Here, we investigate the mineralogical and biochemical properties of the involucrum, as these may hold clues to the aquatic adaptations of cetaceans. We compared bioapatites (enamel, dentine, cementum, and skeletal bone) of cetaceans with those of terrestrial artiodactyls and pachyosteosclerotic ribs of manatees (Sirenia). We investigated organic, carbonate, and mineral composition as well as crystal size and crystallinity index. In all studied variables, bioapatites of the cetacean involucrum were intermediate in composition and structure between those of tooth enamel on the one hand and those of dentine, cementum, and skeletal bone on the other. We also studied the amino acid composition of the cetacean involucrum relative to that of other skeletal bone. The central involucrum had low glycine and hydroxyproline concentrations but high concentrations of nonessential amino acids, unlike most bone samples but similar to the tympanic of hippos and the (pachyosteosclerotic) ribs of manatees. These amino acid results are evidence of rapid bone development. We hypothesize that the mineralogical and amino acid composition of cetacean bullae differs from that of other bone because of (1) functional modifications for underwater sound reception and (2) structural adaptations related to rapid ossification.
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spelling curtin-20.500.11937-110902018-03-29T09:05:57Z Unique Biochemical and Mineral Composition of Whale Ear Bones Kim, S. Thewissen, J. Churchill, M. Suydam, R. Ketten, Darlene Clementz, M. aquatic mammals ear bone cetaceans underwater hearing terrestrial artiodactyls Whales Cetaceans are obligate aquatic mammals derived from terrestrial artiodactyls. The defining characteristic of cetaceans is a thick and dense lip (pachyosteosclerotic involucrum) of an ear bone (the tympanic). This unique feature is absent in modern terrestrial artiodactyls and is suggested to be important in underwater hearing. Here, we investigate the mineralogical and biochemical properties of the involucrum, as these may hold clues to the aquatic adaptations of cetaceans. We compared bioapatites (enamel, dentine, cementum, and skeletal bone) of cetaceans with those of terrestrial artiodactyls and pachyosteosclerotic ribs of manatees (Sirenia). We investigated organic, carbonate, and mineral composition as well as crystal size and crystallinity index. In all studied variables, bioapatites of the cetacean involucrum were intermediate in composition and structure between those of tooth enamel on the one hand and those of dentine, cementum, and skeletal bone on the other. We also studied the amino acid composition of the cetacean involucrum relative to that of other skeletal bone. The central involucrum had low glycine and hydroxyproline concentrations but high concentrations of nonessential amino acids, unlike most bone samples but similar to the tympanic of hippos and the (pachyosteosclerotic) ribs of manatees. These amino acid results are evidence of rapid bone development. We hypothesize that the mineralogical and amino acid composition of cetacean bullae differs from that of other bone because of (1) functional modifications for underwater sound reception and (2) structural adaptations related to rapid ossification. 2014 Journal Article http://hdl.handle.net/20.500.11937/11090 10.1086/676309 The University of Chicago Press restricted
spellingShingle aquatic mammals
ear bone
cetaceans
underwater hearing
terrestrial artiodactyls
Whales
Kim, S.
Thewissen, J.
Churchill, M.
Suydam, R.
Ketten, Darlene
Clementz, M.
Unique Biochemical and Mineral Composition of Whale Ear Bones
title Unique Biochemical and Mineral Composition of Whale Ear Bones
title_full Unique Biochemical and Mineral Composition of Whale Ear Bones
title_fullStr Unique Biochemical and Mineral Composition of Whale Ear Bones
title_full_unstemmed Unique Biochemical and Mineral Composition of Whale Ear Bones
title_short Unique Biochemical and Mineral Composition of Whale Ear Bones
title_sort unique biochemical and mineral composition of whale ear bones
topic aquatic mammals
ear bone
cetaceans
underwater hearing
terrestrial artiodactyls
Whales
url http://hdl.handle.net/20.500.11937/11090